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PHRYMACEAE LOPSEED FAMILY

David M. Thompson

Annual to shrub. Stem: ×-section generally round. Leaf: opposite, simple, entire or toothed; stipules 0. Inflorescence: spike, raceme, or panicle, bracted, or flowers 1–2 in axils. Flower: bisexual; calyx radial or ± bilateral, tube long, generally ribbed, lobes 5; corolla generally bilateral, generally 2-lipped, upper [1]2-lobed, lower 3-lobed; stamens 4 in 2 pairs, epipetalous, included or exserted; pistil 1, ovary superior, chambers 1–2, placentas parietal or axile, style 1, stigma lobes 2, flat, folding together when touched. Fruit: generally capsule, generally ellipsoid, loculicidal [indehiscent, 1-seeded].
± 15 genera, 230 species: ± worldwide, many habitats; some cultivated as ornamental (e.g., Mimulus, Mazus). Formerly included in Scrophulariaceae. [Beardsley & Olmstead 2002 Amer J Bot 89:1093–1102] Family description by David J. Keil. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

MIMULUS MONKEYFLOWER
Glabrous to hairy. Stem: generally erect. Leaf: opposite, generally ± sessile, generally toothed or entire, generally green or ± red. Inflorescence: raceme, bracted, or flowers generally 2 per axil. Flower: occasionally cleistogamous; calyx generally green, lobes 5, generally << tube, generally unequal; corolla generally deciduous, white to red, maroon, purple, gold, or yellow, lower lip base occasionally swollen, ± closing mouth, tube-throat floor generally with 2 longitudinal folds; anther sacs spreading; placentas 2, axile or parietal; stigma lobes generally flat, generally included. Fruit: generally ovoid to fusiform, generally upcurved if elongate, generally ± fragile, loculicidal near tip (hard, indehiscent), chambers 1–2. Seed: many, generally < 1 mm, ovoid, ± yellow to dark brown.
± 100 species: North America, Chile, eastern Asia, southern Africa, New Zealand, Australia. (Latin: little mime or comic actor, from face-like corolla of some species) [Thompson 2005 Syst Bot Monogr 75:1–213] Limb width measured between most distant points across corolla face.

Key to Mimulus

M. aurantiacus Curtis
NATIVE
Subshrub, shrub, glabrous to hairy. Stem: 10–150 cm; main leaf axils generally with small leaf clusters. Leaf: 10–88 mm; upper surface glabrous, lower generally sticky. Inflorescence: pedicel 3–30 mm. Flower: calyx 17–46 mm, not swollen at base, glabrous to hairy, lobes unequal, 2–11 mm, acute to acuminate; corolla persistent, ± yellow, orange, or red, tube-throat 25–57 mm; placentas parietal. Fruit: 13–25 mm, splitting at upper suture.
2n=20. Complex; varieties intergrading, hybridizing. Mimulus aurantiacus subsp. australis Munz, Mimulus aurantiacus subsp. lompocensis (McMinn) Munz are intervarietal hybrids. [Online Interchange]

M. aurantiacus var. pubescens (Torr.) D.M. Thomps.
NATIVE
Leaf: narrowly elliptic, serrate, edges generally ± flat, rolled under in drought; lower surface hairy. Inflorescence: flower 2 per node; pedicel 3–12 mm. Flower: calyx 25–45 mm, tube puberulent to hairy, distally appressed to corolla tube; corolla pale yellow, tube-throat 37–52 mm, lobes entire to 2-lobed; anthers included.
Well-drained, exposed sites; crevices in boulders or rock outcrops in desert areas; 7–2440 m. s Sierra Nevada, s Central Western California, Southwestern California (including Santa Cruz, Santa Rosa islands), s Desert Mountains; northern Baja California. Mar–Jun [Online Interchange]
Unabridged synonyms: [Diplacus arachnoideus Greene; Diplacus calycinus Eastw.; Diplacus speciosus Burtt Davy; Mimulus glutinosus J.C. Wendl. var. brachypus A. Gray; Mimulus longiflorus (Nutt.) A.L. Grant; Mimulus longiflorus var. rutilus A.L. Grant.]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 1 2014
Citation for this treatment: [Author of taxon treatment] 2013. Mimulus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=76756, accessed on Oct 1 2014

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click for enlargement Mimulus aurantiacus var. pubescens
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© 2012 Neal Kramer

Geographic subdivisions indicated for the distribution of Mimulus aurantiacus var. pubescens Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.