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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery.
Leaf: generally simple, alternate; generally both basal, cauline; stipules 0.
Inflorescence: generally raceme, generally not bracted.
Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed.
Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not.
Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Medit area, mtns of sw Asia, adjacent c Asia, w North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in c NCoR, Carrichtera annua (L.) DC. in SCo, Iberis sempervirens L., Iberis umbellata L. in PR, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in s NCoRO, CCo. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; CA taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

THYSANOCARPUS LACEPOD, FRINGEPOD
Annual; hairs 0 or simple.
Leaf: simple, sessile, entire, dentate, or pinnately lobed; middle, distal cauline clasping, lobed or not.
Inflorescence: open ( dense).
Flower: sepals ascending, base not sac-like; petals ± >= sepals, white or ± purple-tinged, not clawed.
Fruit: silicle, indehiscent, elliptic to ovate or round, flat, unsegmented, pendulous; septum 0; wing entire or lobed, generally perforated, veins radiating or not; stigma entire.
Seed: 1, elliptic to round, wingless.
5 species: w North America. (Greek: fringe fruit) Revised taxonomy, too late for full treatment here, including Thysanocarpus desertorum A. Heller [Thysanocarpus laciniatus var. hitchcockii], Thysanocarpus rigidus (Munz) P.J. Alexander & Windham [Thysanocarpus laciniatus var. rigidus], 5 subspp. of Thysanocarpus curvipes (Alexander et al. 2010 Syst Bot 35:559–577).

Key to Thysanocarpus

T. laciniatus Nutt.
NATIVE
Plant generally glaucous, glabrous (sparsely hairy).
Stem: 1–6 dm.
Leaf: 1–6 cm, oblanceolate to elliptic, pinnately lobed, ± entire or occasionally wavy- dentate; mid- cauline linear, tapered at base (minutely lobed, not clasping).
Inflorescence: raceme open.
Fruit: 2.5–5 mm wide, obovate to round, glabrous or occasionally with club-shaped hairs; wing flat or ± curved to convex side, entire to deeply crenate, radiating veins 0 or obscure; pedicels spreading, straight to ± recurved, 3–6(10) mm. Highly variable, probably hybridizes with Thysanocarpus curvipes, resembles the latter when cauline leaf bases lobed. [Online Interchange]

T. laciniatus var. laciniatus
NATIVE

Leaf: ± green or occasionally ± purple, basal pinnately lobed or occasionally wavy- dentate.
Fruit: glabrous or occasionally hairs 0.2–0.4 mm, club-shaped; pedicel ± recurved (± straight).
Oak woodland, rocky ridges, slopes, chaparral, washes; 100–1800 m. Inner North Coast Ranges, s Sierra Nevada Foothills, Tehachapi Mountain Area, Sacramento Valley (Sutter Buttes), San Francisco Bay Area, South Coast Ranges, Southwestern California, East of Sierra Nevada, Desert; w Arizona, Baja California. Mar–May [Online Interchange]
Unabridged note: There are Consortium records that, if verified, would voucher elevations from 3 m to 2163 m. The following (and possibly other) accessions, if verified, would represent range extensions (as indicated): CHSC89031, CHSC89347, UC1713739 (CaRF).

Previous taxon: Thysanocarpus laciniatus var. hitchcockii
Next taxon: Thysanocarpus laciniatus var. rigidus

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
View all CCH records

 

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.