Directory       News       Site Map       Home
         
    Jepson eFlora: Taxon page
Key to families | Table of families and genera

Previous taxon Indexes to all accepted names and synonyms:
| A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |
Previous taxon

ASTERACEAE (Compositae) SUNFLOWER FAMILY

David J. Keil, except as noted

Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1–many in generally ± cyme-like cluster; each head generally with ± calyx-like involucre of 1–many series of phyllaries (involucral bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1–many per head. Flower: bisexual, unisexual, or sterile, ± small, of several types (see below); calyx 0 or modified into ± persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3–5; stamens 4–5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally ± 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing ± brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
± 1500 genera, 23000 species: worldwide, many habitats. Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Bahia; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Taxa of Arida in TJM2 treated here in Leucosyris. —Scientific Editors: David J. Keil, Bruce G. Baldwin.
Unabridged note: Largest family of vascular plants in California and of eudicots globally.

Key to Asteraceae

GRINDELIA GUMPLANT

Abigail J. Moore

[Annual] perennial herb to subshrub from taproot or woody caudex, glabrous or tomentose, often glandular-sticky. Leaf: simple, alternate, generally not fleshy, entire, crenate, serrate, or pinnately lobed, gland-dotted. Inflorescence: heads generally radiate (discoid); involucre obconic to hemispheric, generally gummy; phyllaries in 4–10 graduated series; receptacle flat to convex, ± pitted, epaleate. Ray flower: 0–60; corolla yellow. Disk flower: corolla yellow; anther tip lanceolate; style-branch appendages linear to lanceolate, generally >= stigmatic portion. Fruit: cylindric or swollen-obconic, shiny-white to ± brown, smooth or ridged, glabrous; pappus of 1–6 narrow awns (occasionally construed as bristle-like) [25–40 bristles], ± < disk corolla, generally entire, deciduous.
± 60 species: central and western North America, South America. (D.H. Grindel, Latvian botanist, 1776–1836) [Strother & Wetter 2006 FNANM 20:424–436] Variable. Morphologically intermediate plants common where species ranges overlap. Strother & Wetter treated all California species except Grindelia squarrosa and Grindelia fraxinipratensis in Grindelia hirsutula. Grindelia ciliata (Nutt.) Spreng. [Prionopsis ciliata (Nutt.) Nutt.], with pappus of many united bristles (falling as a unit), is an historical waif from northern Central Coast, northern South Coast.

Key to Grindelia

G. squarrosa (Pursh) Dunal var. serrulata (Rydb.) Steyerm.
NATURALIZED
Biennial 1–6 dm, decumbent to erect, much-branched throughout. Leaf: 1.5–7 cm; basal leaves generally 0 at flower, distal not much smaller; blade oblong to ovate, sessile or narrowed at base, glabrous, resinous, gray-green, crenate, each tooth with a ± yellow bump near tip. Inflorescence: involucre 10–17 mm diam, bell-shaped, glabrous, resinous; phyllaries in 5–6 series, bases wide, straw-colored, tips green, acuminate, ± round in ×-section, coiled 360°. Ray flower: 0 or 24–36; ray 8–10 mm. Fruit: 2.3–3 mm, light brown to ± yellow, top truncate; pappus awns 2–3(6).
2n=12. Disturbed roadsides, streamsides; 700–2300 m. High Cascade Range, High Sierra Nevada, Transverse Ranges, Great Basin Floristic Province, Mojave Desert; native Wyoming to New Mexico. TOXIC, concentrates selenium. Jul–Sep [Online Interchange]

Previous taxon: Grindelia X paludosa
Next taxon: Grindelia stricta

Contact/Feedback

Name search

Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 23 2014
Citation for this treatment: [Author of taxon treatment] 2013. Grindelia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=7515, accessed on Nov 23 2014

Copyright © 2014 Regents of the University of California
We encourage links to these pages, but the content may not be downloaded for reposting, repackaging, redistributing, or sale in any form, without written permission from The Jepson Herbarium.


Geographic subdivisions indicated for the distribution of Grindelia squarrosa var. serrulata Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
READ ABOUT YELLOW FLAGS
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

View elevation by latitude chart
Data provided by the participants of the Consortium of California Herbaria.
View all CCH records

 

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.