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Key to families | Table of families and genera
Indexes to all accepted names and synonyms:
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Annual, perennial herb generally from rhizomes. Stem: round or flat. Leaf: generally basal; sheath margins fused, or overlapping and generally with 2 ear-like extensions at blade junction; blade round, flat, or vestigial, glabrous or margin hairy. Inflorescence: head-like clusters or flowers 1, variously arranged; bracts subtending inflorescence 2, generally leaf-like; bracts subtending inflorescence branches 1–2, reduced; bractlets subtending flowers generally 1–2, generally translucent. Flower: generally bisexual, radial; sepals and petals similar, persistent, scale-like, green to brown or ± purple-black; stamens generally 3 or 6, anthers linear, persistent; pistil 1, ovary superior, chambers generally 1 or 3, placentas 1 and basal or 3 and axile or parietal, stigmas generally > style. Fruit: capsule, loculicidal. Seed: 3–many, generally with white appendages on 1 or both ends.
7 genera, 440 species: temperate, arctic, and tropical mountains. [Kirschner 2002 Species Plantarum: Fl World, vols. 6–8 (Juncaceae). ABRS] Flowers late spring to early fall. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Balslev, H. 1996. Juncaceae. Flora Neotropica Monograph 68: 1–168. New York Botanical Garden, Bronx, NY; —Brooks, R. E. and S.E. Clemants. 2000. Juncus. In: Flora of North America Editorial Committee, Eds. Flora of North America North of Mexico. Volume 22, Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in part), and Zingiberidae. Oxford University Press, New York. Pages 211–255; —Brooks, R. E. and A.T. Whittemore. 1999. Juncus anthelatus (Juncaceae, Juncus subg. Poiophylli), a new status for a North American taxon. Novon 9:11–12; —Catling, P. M. and K.W. Spicer. 1987. The perennial Juncus of section Poiophylli in the Canadian prairie provinces. Canadian Journal of Botany 65: 750–760; —Ceska A. 2001. Juncaceae. In: Douglas, G. W., D. Meidinger, and J. Pojar, Eds. Illustrated Flora of British Columbia, Volume 6, Monocotyledons (Acoraceae Through Najadaceae). British Columbia Ministry of Environment, Lands and Parks, Ministry of Forests, Victoria, British Columbia, Canada; —Clemants, S. E. 1990. Juncaceae (Rush Family) of New York State. New York State Museum Bulletin No. 475, Albany, NY; —Cope, T. A. and C.A. Stace. 1978. The Juncus bufonius L. aggregate in western Europe. Watsonia 12: 113–128; —Drábková, L., J. Kirschner, O. Seberg, G. Petersen and C. Vlcek. 2003. Phylogeny of the Juncaceae based on rbcL sequences, with special emphasis in Luzula DC. and Juncus L. Plant Systematics and Evolution 240: 133–147; —Drábková, L., J. Kirschner and C. Vlcek. 2002. Comparison of seven DNA extraction and amplification protocols in historical herbarium specimens of Juncaceae. Plant Molecular Biology Reporter 20: 161–175; —Drábková, L., J. Kirschner, C. Vlcek and V. Paces. 2004. TrnL-trnF intergenic spacer and trnL intron define major clades within effusus aggregateuzula and Juncus (Juncaceae): importance of structural mutations. Journal of Molecular Evolution 59: 1–10; —Ertter, B. 1986. The Juncus triformis complex. Memoirs of the New York Botanical Garden 39: 1–89; —Hämet-Ahti, L. 1980. The Juncus effusus aggregate in eastern North America. Annales Botanici Fennici 17: 183–191; —Harriman, N. A. and D. Redmond. 1976. Somatic chromosome numbers for some North American species of Juncus L. Rhodora 78: 727–738; —Hermann F. J. 1964. The Juncus mertensianus complex in western North America. Leaflets of Western Botany 10: 81–87; —Hermann F. J. 1975. Manual of the Rushes (Juncus species) of the Rocky Mountains and Colorado Basin. USDA Forest Service, General Technical Report RM-18, Fort Collins, CO; —Johnson, L. A. S. 1963. New species of Juncus in Australia and New Zealand. Contributions from the New South Wales National Herbarium 3: 241–244; —Jokerst, J. 1986. A new Juncus from Chile. Fremontia 14(2): 22–23; —Kirschner, J., Ed. 2002. Juncaceae 1: Rostkovia to Luzula, Species Plantarum: Flora of the World Part 6. Australian Biological Resources Study, Canberra, Australia. 237 p; —Kirschner, J., Ed. 2002. Juncaceae 2: Juncus subg. Juncus, Species Plantarum: Flora of the World Part 7. Australian Biological Resources Study, Canberra, Australia. 336 p; —Kirschner, J., Ed. 2002. Juncaceae 3: Juncus subg. Agathryon, Species Plantarum: Flora of the World Part 8. Australian Biological Resources Study, Canberra, Australia. 192 p; —Lamont, E. E. and S.M. Young. 2005. Juncus diffusissimus, an addition to the flora of New York, with notes on its recent spread in the United States. Journal of the Torrey Botanical Society 132: 635–643; —Lint, H. L. 1977. A revision of Juncus subgenus Genuini (Juncaceae) in the Pacific States. Unpublished Ph.D. Dissertation, Oregon State University, Corvallis; — Munro, S. L. and H.P. Linder. 1998. The phylogenetic position of Prionium (Juncaceae) within the Order Juncales based on morphological and rbcL sequence data. Syst Bot 23: 43–55; —Plunkett, G. M., D.E. Soltis, P.S. Soltis, and R.E. Brooks. 1995. Phylogenetic relationships between Juncaceae and Cyperaceae: insights from rbcL sequence data. 1995. American Journal of Botany 82: 520–525; —Roalson, E. H. 2005. Phylogenetic relationships in the Juncaceae inferred from nuclear ribosomal DNA internal transcribed spacer sequence data. International Journal of Plant Sciences 166: 397–413; —Snogerup, S. 1978. Notes on Juncus for Flora Europaea. Botaniska Notiser 131: 185–187; —Snogerup, S. 1980. Juncaceae. Pp. 102–116. In: Tutin, T. G., V.H. Heywood, N.A. Burges, D.M. Moore, D.H. Valentine, S.M. Walters and D.A. Webb, eds. 1980. Flora Europaea, Vol. 5, Alismataceae to Orchidaceae (Monocotyledones). Cambridge Univ. Press, Cambridge, UK. 452 p; —Snogerup, S. 1993. A revision of Juncus subgen. Juncus (Juncaceae). Willdenowia 23: 23–73; —Snogerup, S., P.F. Zika, and J. Kirschner. 2002. Taxonomic and nomenclatural notes on Juncus. Preslia 74: 247–266. [Available at: http://www.ibot.cas.cz/preslia/P023CSno.pdf] Stace, C. A. 1970. Anatomy and taxonomy in Juncus subgenus Genuini. Botanical Journal of the Linnean Society 63 (suppl. 1): 75–84; —Stuckey, R. L. 1980. The migration and establishment of Juncus gerardii (Juncaceae) in the interior of North America. Sida 8: 334–347; —Van Loenhoud, P. J. and A.A. Sterk. 1976. A study of Juncus bufonius complex in the Netherlands. Acta Botanica Neerlandica 25: 193–204; —Witham, C. W. and P.F. Zika. 2008. Juncus digitatus (Juncaceae), a new annual rush from Shasta County, California, U.S.A. Journal of the Botanical Research Institute of Texas 2: 775–781; —Zika, P. F. 2003. The native subspecies of Juncus effusus (Juncaceae) in western North America. Brittonia 55: 150–156; —Zika, P. F. 2006. A key to Juncus section Juncotypus in British Columbia. Botanical Electronic News No. 358, February 27, 2006. [Available at: http://www.ou.edu/cas/botany-micro/ben/ben358.html#2]; Zika, P. F. 2012. Juncus trilocularis (Juncaceae), a new rush species from western North America. Rhodora 114: 309–329.]
Key to Juncaceae
Perennial herb, cespitose or rhizomatous, rhizome ascending or vertical. Stem: cylindric. Leaf: generally basal, cauline few; blades linear, flat or channeled, margins and sheath opening, sparsely to densely long-soft-hairy. Inflorescence: panicles of 1–few flowers per branch, or head-like and cylindric to ovoid in dense to loose clusters; lower bract leaf-like or herbaceous at base, membranous distally, bractlets 1–3, margins generally ciliate. Flower: perianth parts 6, pale brown to ± black-brown; stamens 6; pistil 1, chamber 1, placenta basal. Fruit: capsule, opening with 3 valves. Seed: 3, elliptic to ovoid, ridged on 1 side, occasionally attached to placenta by tuft of hairs, generally with ± white appendage at tip.Key to Luzula
± 115 species: worldwide. (Latin: light; Italian: glow worm) Measure unridged side of seeds; seed length measurement not including appendage.
Cespitose, < 60 cm, rhizome short; stolons 0. Leaf: flat, ciliate, tip obtuse, thickened. Inflorescence: peduncled to congested, elongated clusters with basal flowers generally remote, lower part of clusters generally loose; lower bract herbaceous. Flower: perianth parts 2.5–4 mm, ± equal, lanceolate, acuminate, pale straw-brown to pale brown in age. Fruit: ± < perianth. Seed: seeds ovoid to narrowly ovoid; appendage 0.4–0.5 mm.
2n=12,24. Occasionally misidentified as Luzula subsessilis. Plants from San Gabriel Mountains, San Bernardino Mountains, San Jacinto Mountains need further study. [Online Interchange]
Unabridged synonyms: [Luzula campestris var. comosa (E. Mey.) Fernald & Wiegand]
Unabridged note: Many literature records may belong to Luzula subsessilis, particularly those from the south. Very variable even when a complex of populations of Luzula subsessilis is excluded. It comprises two cytotypes, roughly corresponding to two varieties (nature of intermediates remains unknown).
Plant 25–60 cm. Leaf: basal leaves 8–13 cm, 3.5–6 mm wide; cauline 3–4, 6–10 cm, 2–4.5 mm wide. Inflorescence: densely to loosely congested, 1.5–3 cm, ± 1–1.5 cm wide, ± 8–15-flowered, ± small-lobed, basal cluster generally remote or long-peduncled in axil of upper cauline leaf, distal clusters generally short-peduncled; lower bract 1.5–4.5 cm, > inflorescence. Flower: perianth parts 2.5–3.7 mm, pale straw-brown to ± brown in age; anthers 0.6–1 mm, filaments 0.5–0.6 mm, style 0.4–0.5 mm, stigma ± 1.3–1.8 mm. Fruit: valves ± 2.3–2.8 mm, 1.4–1.8 mm wide. Seed: ovoid, 1–1.2 mm, 0.8–0.9 mm wide; appendage ± 0.5 mm.
2n=24. Meadows, open woodland, conifer forest; generally < 1500 m. Northwestern California, Sierra Nevada, Central Coast, San Francisco Bay Area; to Alaska. [Luzula comosa var. congesta S. Watson] Jun–Jul [Online Interchange]
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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on May 5 2015
Citation for this treatment: [Author of taxon treatment] 2013. Luzula, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=72790, accessed on May 5 2015
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|Geographic subdivisions indicated for the distribution of Luzula comosa var. comosa|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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