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BORAGINACEAE BORAGE or WATERLEAF FAMILY

Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted

Annual to shrub, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1–2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages 0 or 5 at top of tube, alternate stamens; stamens epipetalous; ovary superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1–4, free (fused), smooth to roughened, prickly or bristly or not.
± 120 genera, ± 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum). Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. [Olmstead et al. 2000 Molec Phylogen Evol 16:96–112] Recently treated to include Hydrophyllaceae, Lennoaceae. —Scientific Editors: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.

Key to Boraginaceae

CRYPTANTHA

Ronald B. Kelley, Michael G. Simpson & Kristen E. Hasenstab-Lehman

Annual to perennial herb, generally erect. Stem: branches 0 or generally ascending to erect, hairy. Leaf: generally sessile; basal whorled, cauline generally alternate, reduced above (opposite below); generally strigose, rough-hairy, or bristly, largest bristles generally bulbous-based. Inflorescence: generally terminal, raceme- or generally spike-like cymes, in groups of 1–5 (> 5), generally coiled in bud, generally elongated in fruit; bracts generally 0. Flower: generally unscented, persistent or not; sepals fused at base; corolla tube generally 1–13 mm, limb 0.5–12 mm diam, generally white, appendages generally 5; anthers included; ovary generally 4-lobed. Fruit: pedicel 0 or < 12 mm in fruit; nutlets 1–4, generally gray to brown, smooth to granular, tubercled, or papillate, with abaxial, longitudinal ridge to not; margin rounded, sharp-edged, or a ± flat rim or wing; adaxially grooved above attachment scar, scar generally lateral, narrow, open to closed, raised or generally not, edges inrolled to sharp-angled, generally forked or flared open at base; central fruit axis ("axis") not reaching to extending beyond fruit.
± 200 species: western North America, western South America, northeastern Asia (1 sp.). (Greek: hidden flowers, from cleistogamous flowers of some South American species) Generally homostylous. The tissue between ovary lobes, interpreted as a receptacle and/or style (style sometimes 0, then stigma attached to top of receptacle), extends to various degrees in fruit, forming what is often called the gynobase (here "fruit axis"), to which the nutlets are laterally attached at maturity, leaving an attachment scar. Annual species without yellow corolla appendages generally self-pollinating; perennial herb species generally homostylous in California. Some species, e.g., Cryptantha angustifolia, Cryptantha ambigua, Cryptantha barbigera, Cryptantha mariposae, hybridize with co-occurring species. Observation of nutlets, hairs best at 10+× generally critical for identification. Corolla limb diam generally < at end of flower period, especially for annual species Cryptantha sobolifera Payson does not occur in California.
Unabridged references: [Johnston 1925 Contr Gray Herbarium 74:1–125; Higgins 1971 Brigham Young Sci Bull Biol Ser 13:1–63, 1979 Great Basin Naturalist 39:293–350; Simpson & Hasenstab 2009 Crossosoma 35:1–59]

Key to Cryptantha

C. barbigera (A. Gray) Greene
NATIVE
Annual 6–50 cm, erect, stout, white-hairy. Stem: 1–few, branches 0 generally below middle; sparse-strigose and generally dense-bristly. Leaf: 1–5(8) cm, oblong to ± blunt-lanceolate; bristles dense, spreading, minute-bulbous-based. Inflorescence: in (1s) 2s (3s), generally many-flowered; bracts 0; pedicel 0–0.5 mm, ascending to spreading, generally spreading-soft-hairy. Flower: calyx 3–6 mm, generally constricted above, 1 lobe generally longer, midvein thickened, spreading-bristly. Fruit: nutlets ovate to lance-ovate, dense-tubercled, ± brown, occasionally mottled, dull, margin ± rounded, base truncate, tip tapered to ± beaked; abaxially rounded, ridge 0; adaxially flat, attachment scar edges not raised, abutted entire length or variously gapped, triangular-flare-gapped at base; axis to nutlets or beyond. [Online Interchange]

C. barbigera var. barbigera BEARDED CRYPTANTHA
NATIVE
Flower: calyx 5–10 mm in fruit, lobes generally spreading or recurved, generally lance-linear, margins generally dense-soft-hairy with spreading, fine bristles below middle; corolla deciduous, limb 1–2.5(3) mm diam, appendages 0 or minute, white. Fruit: nutlets 1–4, 1.5–2 mm.
Open, sandy to rocky soils, generally in creosote-bush scrub, desert woodland; 15–1860(2250) m. s Sierra Nevada, Tehachapi Mountain Area, s San Joaquin Valley, s East of Sierra Nevada, Desert; to Utah, New Mexico, Baja California. Feb–Jun [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 25 2014
Citation for this treatment: [Author of taxon treatment] 2013. Cryptantha, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=71929, accessed on Jul 25 2014

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click for enlargement Cryptantha barbigera var. barbigera
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© 2014 Steve Matson

Bioregions in which Cryptantha barbigera var. barbigera occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.