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ONAGRACEAE EVENING-PRIMROSE FAMILY

Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted

Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2–7); petals 4(2–7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
22 genera, ± 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Unabridged references: [Munz 1965 North America Fl II 5:1–278]

Key to Onagraceae

CHAMERION

Peter C. Hoch

Perennial herb, often clumped or forming large colonies by rhizomes. Stem: generally unbranched, strigose or glabrous. Leaf: alternate, generally ± fine-toothed; veins conspicuous or obscure. Inflorescence: raceme. Flower: nodding in bud; hypanthium 0 (except as ± green disk); ± bilateral; sepals 4, spreading; petals 4, entire; stamens 8, subequal, maturing before stigma, anthers attached at middle, pollen grains shed singly, generally blue-gray; stigma spreading-4-lobed. Fruit: straight, cylindric. Seed: many in 1 row per chamber, irregularly netted, with persistent white hair-tuft.
8 species: northern temperate montane, boreal. [Wagner et al. 2007 Syst Bot Monogr 83:78–81]
Unabridged references: [Raven 1976 Ann Missouri Bot Gard 63:326–340]

Key to Chamerion

Previous taxon: Camissoniopsis robusta
Next taxon: Chamerion angustifolium subsp. circumvagum

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 25 2014
Citation for this treatment: [Author of taxon treatment] 2013. Chamerion, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=68252, accessed on Oct 25 2014

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