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John C. Game, Alan R. Smith & Thomas Lemieux

Plants generally in soil; rhizome short- to long-creeping, prostrate to erect, [trunk-like], scales generally hairy, ± brown. Leaf: stipe ×-section at base with 2 crescent-shaped vascular strands fusing into 1 U-shaped strand distally; blade 1[2]-pinnate, generally with needle-like, clear hairs on axes and/or between veins; rachis, costae generally grooved adaxially except between axes; veins free or in regular, net-like pattern. Sporangia: sori on veins, generally round; indusia reniform or round-reniform [0]; stalk cells in 3 rows; spores generally elliptic, scar generally linear.
1 genus (5–32 genera), ± 1000 species: worldwide, especially tropics; several cultivated. —Scientific Editors: Alan R. Smith, Thomas J. Rosatti.

Leaf: pinnae entire to generally deeply lobed, veins generally not forked. (Greek: female fern; ancient name for delicate fern) [Smith & Cranfill 2002 Amer Fern J 92:131–149]
Unabridged references: [Smith 1971 Univ Calif Publ Bot 59:1–143; Smith, A. R. 1990. Thelypteridaceae. Pp. 263–272 in The families and genera of vascular plants, ed. K. Kubitzki. Vol. 1. Pteridophytes and gymnosperms, ed. K. Kramer & P. Green. Berlin: Springer-Verlag]

Key to Thelypteris

T. puberula (Baker) C.V. Morton var. sonorensis A.R. Sm. SONORAN MAIDEN FERN
Rhizome creeping, 3–8 mm wide; evergreen. Leaf: ± regularly spaced, generally 50–120 cm, 15–30 cm wide; stipe scales lanceolate, brown, not persistent; blade thick, abaxially ± nonglandular, ± densely hairy on axes, veins, and between veins; pinnae deeply lobed, lobes narrowly oblong, entire. Sporangia: sori small, round; indusia densely hairy.
Along streams, seepage areas; 50–800 m. South Coast, Western Transverse Ranges, San Gabriel Mountains, San Jacinto Mountains; to Arizona, southern Mexico. Rarely cultivated in southern California. 1 other var., Thelypteris puberula var. puberula, Mexico to Costa Rica. [Online Interchange] {CNPS list}
Unabridged note: Christella puberula A. Löve & D. Löve, published [1977 Taxon 26:326] for the nominate element of Thelypteris puberula, may reflect a superior classification, but a combination that would include var. sonorensis within Christella puberula has not been made. Recent molecular evidence [A.R. Smith & R.B. Cranfill 2002 Taxon 56:1037–1050] suggests that Christella itself may be polyphyletic and possibly not separable from Cyclosorus s.l. Some combinations of supposed Christella taxa within Cyclosorus, including "Cyclosorus puberulus," have not yet been made.

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 27 2015
Citation for this treatment: [Author of taxon treatment] 2013. Thelypteris, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 27 2015

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Geographic subdivisions indicated for the distribution of Thelypteris puberula var. sonorensis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.