Jepson Herbarium
		University of California, Berkeley

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Annual, perennial herb, woody vine [ shrub], occasionally aquatic.
Leaf: generally basal and cauline, alternate or opposite, simple or compound; petioles at base generally flat, occasionally sheathing or stipule-like.
Inflorescence: cyme, raceme, panicle, or flowers 1.
Flower: generally bisexual, generally radial; sepals 3–6(20), free, early- deciduous or withering in fruit, generally green; petals 0–many, generally free; stamens generally 5–many, staminodes generally 0; pistils 1–many, ovary superior, chamber 1, style 0–1, generally ± persistent as beak, ovules 1–many.
Fruit: achene, follicle, berry, ± utricle in Trautvetteria, in aggregate or not, 1–many-seeded.
± 60 genera, 1700 species: worldwide, especially n temperate, tropical mountains; many ornamental (Adonis, Aquilegia, Clematis, Consolida, Delphinium, Helleborus, Nigella). some highly TOXIC (Aconitum, Actaea, Delphinium, Ranunculus). [Whittemore & Parfitt 1997 FNANM 3:85–271] Taxa of Isopyrum in TJM (1993) moved to Enemion; Kumlienia moved to Ranunculus. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Ranunculaceae

Perennial from caudex or rhizomes, generally glabrous; dioecious or flowers bisexual.
Stem: 1–few, generally erect; branches 0 or few.
Leaf: 1–4- ternate or pinnate, basal or basal and cauline, alternate, generally reduced distally on stem; leaflets wedge-shaped to ± round, entire, crenate, or lobed; pale green abaxially, generally green adaxially.
Inflorescence: raceme or panicle, axillary or terminal, generally erect, ± scapose or not; bracts simple to 1- ternate; pedicels generally erect in fruit.
Flower: sepals 4–5, ± green-white to ± purple, petal-like or not, generally early- deciduous; petals 0; stamens 8–many, generally > sepals, filaments flat or generally thread-like, anthers generally narrowly oblong, tip generally pointed; pistils (1)2–22.
Fruit: achene, compressed laterally or not, ribbed or veined, beaked.
120–200 species: temperate North America, Eurasia, Africa; some ornamental, medicinal. (Greek: name given by Dioscorides, Greek physician-botanist) [Park & Festerling 1997 FNANM 3:258–271]

Plant 60–200 cm; generally dioecious.
Leaf: basal and cauline, 7–46 cm; segments 8–20 mm, glabrous to finely glandular-puberulent.
Inflorescence: panicle, bracts leaf-like proximally.
Flower: sepals generally 4, 2–5 mm; stamens 15–28, filaments thread-like, anthers oblong- linear, acuminate.
Fruit: 7–20, spreading to ascending; body 4–8 mm, ± compressed laterally, sides obliquely ± ovate or ± widely obovate to ± round, ribs 1–3, ± curved, veins 0–several, beak 1.5–4 mm. Plants in NCoR occasionally with bisexual flowers; vars. in CA difficult, need study. [Online Interchange]
Unabridged note: Expanded author citation: Thalictrum fendleri Engelm. ex A. Gray

Leaf: distal abaxially generally glabrous.
Fruit: body compressed generally only near margins, sides obliquely ± widely obovate to ± round, generally 1-ribbed, veins several, wavy, net-like.
2n=28. Moist, open to shaded places, woodland, forest; < 2600 m. North Coast, w Klamath Ranges, North Coast Ranges, Sierra Nevada (± rare), Central Western California, Transverse Ranges, w Peninsular Ranges;

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.