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David J. Keil, except as noted

Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1–many in generally ± cyme-like cluster; each head generally with ± calyx-like involucre of 1–many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1–many per head. Flower: bisexual, unisexual, or sterile, ± small, of several types (see below); calyx 0 or modified into ± persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3–5; stamens 4–5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally ± 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing ± brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
± 1500 genera, 23000 species: worldwide, many habitats. Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Bahia; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Taxa of Arida in TJM2 (2012) treated here in Leucosyris. —Scientific Editors: David J. Keil, Bruce G. Baldwin.
Unabridged note: Largest family of vascular plants in California and of eudicots globally.

Key to Asteraceae


James D. Morefield

Annual 0.5–10(20 cm), ± green to gray, woolly to cobwebby. Stem: 0, or 1, erect, or 2–10, ± decumbent to ascending, generally unbranched or branched proximally, equally or more leafy distally. Leaf: alternate, sessile or petioled, oblanceolate to spoon-shaped, entire, base ± yellow, ± hard; distal leaves subtending heads, ± crowded, largest >= proximal leaves. Inflorescence: heads disciform, sessile, single or 2–40+ per group; involucre 0, simulated by paleae or leaves; receptacle length 0.8–1.3 or 4–6 × width, ± conic, acuminate, bristly; paleae of pistillate flowers open, lanceolate to obovate, acute, flat to concave or ± folded, persistent, ± glabrous, obscurely parallel-veined, veins 5+, margin ± continuous, thinned but scarcely winged; paleae of disk flowers generally 5, ± enlarged, erect proximally, erect to spreading distally, open, obovate, obtuse, cartilaginous, proximally ± yellow, distally green and tomentose adaxially. Pistillate flower: (3)5–25, all subtended by paleae; corolla obscure, narrowly cylindric. Disk flower: staminate, 2–6(12); pappus 0; corolla (3)4(5)-lobed, ± bilateral; anther base tailed, tip ± triangular; style tips ± linear-oblong; pappus 0. Fruit: falling free of paleae, ± obovoid, compressed front-to-back, smooth, dull to ± shiny, corolla scar terminal; pappus 0.
3 species: western California, southern Oregon. (Greek: western Evax) [Morefield 2006 FNANM 19:467–470] Apparent sister of Ancistrocarphus; bristly receptacle unique. Historical California specimens of Diaperia [Evax sect. Diaperia], native central Arizona and eastward, are likely label errors.
Unabridged references: [Morefield 1992 Syst Bot 17:293–310]

Key to Hesperevax

H. acaulis (Kellogg) Greene DWARF EVAX
Stem: 0 or 1–10, prostrate to erect, to 7 cm. Leaf: distal > proximal, longest 4–22(32) mm, < 5 mm wide; petiole 0–1.5 × blade, base scarcely thickened. Inflorescence: distal heads generally 1, terminal, 2–4 mm, 1.5–3.5 mm wide, ± bell-shaped, length 1–1.5 × width, subtended by generally 6–12 leaves (in varieties acaulis and robustior distal heads rarely 2–8 per group, largest group dense, 3–7 mm wide, mixed with leaves); proximal heads 0 or axillary, < distal; receptacle length 1–1.3 × width; paleae in spiral ranks; paleae of disk flowers 0.6–0.8 × head, tips exserted, spreading. Disk flower: 0.6–1 mm. Fruit: 0.6–1.6 mm. Varieties intergrade. [Online Interchange]
Unabridged note: Butte, Tehama cos. reports of Hesperevax acaulis var. acaulis, Hesperevax acaulis var. ambusticola await confirmation.

H. acaulis var. robustior Morefield
Plants generally 2–7 cm. Stem: 1(7), ± erect; branches generally 0. Leaf: largest (9)12–22(32) mm, 2–4(5) mm wide, distal erect or tips spreading; petiole 0–0.8 × blade; blade oblanceolate to obovate, acute to obtuse. Inflorescence: distal heads 3–4 mm, 2.5–3.5 mm wide; receptacle 1.4–1.9 mm; paleae of pistillate flowers generally in 3–5 series, 2.5–4 mm; longest paleae of disk flowers 2.5–3.2 mm. Fruit: generally 1–1.6 mm.
Slopes, flats, swales, canyons, path edges, openings or under shrubs; 60–1100 m. Klamath Ranges, Outer North Coast Ranges, Inner North Coast Ranges, Cascade Range Foothills, Sierra Nevada Foothills, n&e Great Central Valley, Outer South Coast Ranges; to southwestern Oregon. Largest sizes above from cultivation specimen. Apr–Jun [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 28 2015
Citation for this treatment: [Author of taxon treatment] 2013. Hesperevax, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 28 2015

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Geographic subdivisions indicated for the distribution of Hesperevax acaulis var. robustior Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.