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AMARANTHACEAE AMARANTH FAMILY

Mihai Costea, except as noted

Annual to subshrub; monoecious and/or dioecious; occasionally spiny; hairs simple (branched). Leaf: blade simple, alternate or opposite, margins entire or serrate; veins pinnate; stipules 0. Inflorescence: axillary or terminal; 3-flowered cymes in dense spikes, heads or panicles; bracts 0 or 1–5, persistent; bractlets 0–2. Flower: bisexual or unisexual, small, green (± white), yellow or purple; perianth parts 0 or (1)3–5, free or fused basally, scarious or hardened, persistent; stamens 1–5, opposite perianth parts, free or basally fused as a tube, generally unequal, occasionally alternate with appendages on stamen tubes (pseudostaminodes), anthers 2- or 4-chambered; ovary superior, chamber 1; ovule 1 (2–many); style (0)1–3, stigmas 1–3(5). Fruit: utricle; generally with persistent perianth or bracts. Seed: 1 [2+], small, lenticular to spheric, smooth or dotted to striate or tubercled.
± 75 genera, 900 species: cosmopolitan, especially disturbed, arid, saline or alkaline soils; some cultivated for food, ornamental; many naturalized, ruderal or agricultural weeds. [Müller & Borsch 2005 Ann Missouri Bot Gard 92:66–102] Amaranthaceae including Chenopodiaceae by some. Polycnemoideae, represented in California by Nitrophila, formerly considered subfamily of Chenopodiaceae, but needs further research. Guilleminea densa (Willd.) Moq. var. aggregata Uline & W.L. Bray is a waif. Froelichia gracilis (Hook.) Moq. is an historical waif. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Kadereit et al. 2003 Int J Plant Sci 164:959–986; Robertson 1981 J Arnold Arbor 62:267–313]

Key to Amaranthaceae

GUILLEMINEA
Perennial herb, generally prostrate, soft-hairy to woolly, hairs simple. Leaf: opposite; petioled; blade linear or lanceolate to spoon-shaped or ovate, margins entire. Inflorescence: axillary, sessile, few-flowered heads; bracts, bractlets membranous, white, glabrous, persistent. Flower: bisexual; perianth bell-shaped, perianth parts 5, fused ± 1/2, 1-veined; stamens 5, filaments fused basally into tube attached to top of perianth tube, anthers 2-chambered, pseudostaminodes 0; ovary ovoid, ovule 1, style 1, ± 0.2 mm, stigmas 2-lobed, spheric. Fruit: utricle, wall membranous. Seed: 1 lenticular to spheric, smooth, shiny.
± 5 species: southern United States to South America. (Antoine Guillemin, French botanist, explorer, 1796–1842) [Henrickson 1987 Sida 12:307–337]

G. densa (Willd.) Moq. var. aggregata Uline & W.L. Bray SMALL MATWEED
WAIF
Mats 10–80 cm wide. Stem: ± densely woolly. Leaf: basal in rosettes, withering, petiole winged, 8–20 mm, blade(8)20–40 mm, (1)6–9(13) mm wide, oblong-elliptic to lanceolate, glabrous adaxially, woolly abaxially; cauline petiole 1–5 mm, blade 20–100(220) mm, 20–80(150) mm wide, ovate, oblong-ovate, oblong, rhombic, or occasionally elliptic to lanceolate, ± glabrous adaxially, woolly abaxially. Inflorescence: axillary clusters of heads 2–3 mm, 3–5 mm wide, white; bracts 1.5–2.5 mm, white-scarious. Flower: perianth 0.8–2.8 mm, tube 0.8–1.2 mm, lobes reflexed, erect in age, ovate to elliptic, tip acute to rounded, shaggy-hairy especially near tube base. Fruit: included within persistent perianth, 1.1–1.4 mm. Seed: 0.8–0.9 mm, red-brown.
Roadsides, disturbed areas; < 100 m. s Central Coast, South Coast; native Arizona to Texas, South America. Unabridged synonyms: [Brayulinea densa (Willd.) Small]
Aug–Nov [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 16 2014
Citation for this treatment: [Author of taxon treatment] 2012. Guilleminea, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=59691, accessed on Apr 16 2014

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Bioregions in which Guilleminea densa var. aggregata occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.