Jepson Herbarium
		University of California, Berkeley

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Indexes to all accepted names and synonyms: | A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |

Annual to shrub, or non-green root parasite, generally bristly or sharp-hairy.
Stem: prostrate to erect.
Leaf: cauline, often with basal rosette, simple or compound, generally alternate.
Inflorescence: cymes, generally elongate, panicle-, raceme-, or spike-like, generally coiled in flower (often described as scorpioid), generally uncoiled in fruit, or heads, spikes, or panicles, or flowers 1–2 per axil.
Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla generally (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages 0 or 5 at top of tube, alternate stamens; stamens epipetalous; ovary superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched.
Fruit: nutlets 1–4, free ( fused), smooth to roughened, prickly or bristly or not, or valvate or circumscissile capsule.
± 120 genera, ± 2300 species: tropics, temperate, especially w North America, Medit; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum). Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. [Olmsted et al. 2000 Molec Phylogen Evol 16:96–112] Recently treated to include Hydrophyllaceae, Lennoaceae. —Scientific Editors: Ronald B. Kelley, Robert Patterson, Bruce G. Baldwin, Thomas J. Rosatti.

Key to Boraginaceae

Perennial to shrub.
Stem: prostrate to ascending or erect; bark shredding.
Leaf: simple, cauline, alternate.
Inflorescence: generally open, terminal.
Flower: corolla funnel- to urn-shaped, white, lavender, or purple, generally hairy abaxially; stamens included, filaments generally hairy; ovary chambers 2, styles 2, generally hairy.
Fruit: 1–3 mm wide; valves 4.
Seed: striate, dark brown or black.
11 species: sw US, Mex. (Greek: wool net, from abaxial leaves) [Ferguson 1998 Syst Bot 23:253–268]
Unabridged etymology: (Greek: erio, wool, plus dictyon, net, from abaxial leaves)
Unabridged references: [Hannan 1988 Amer J Bot 75:579–588]

Stem: 1–3 m; twigs dense-tomentose.
Leaf: petiole 3–15 mm; blade 3–17 cm, 10–40 mm wide, lanceolate to ovate, ± entire to coarse-toothed, adaxially sparse-hairy to dense-tomentose, abaxially tomentose.
Flower: calyx lobes 2–4 mm, dense-long-hairy; corolla 6–16 mm, funnel-shaped, lavender, abaxially dense-hairy; styles 3–8 mm.
Seed: 8–14.
n=14. Varieties intergrade, ± distinguished by stem, leaf hairs. [Online Interchange]

Stem: dense-gray- to -white tomentose, dark stem obscured.
Leaf: ± ovate, dense-gray- to -white- tomentose.
Flower: corolla 8–16 mm.
Slopes, roadsides, washes, river bottoms, mesas, chaparral, woodland; 15–1520 m. South Coast, Western Transverse Ranges, San Gabriel Mountains, Peninsular Ranges.

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Next taxon: Eriodictyon crassifolium var. nigrescens

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.