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Key to families | Table of families and genera
Indexes to all accepted names and synonyms:
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Annual, perennial herb, shrub, tree, often vine; sap generally milky. Leaf: simple, alternate, opposite, subwhorled to whorled, entire; stipules 0 or small, finger-like. Inflorescence: axillary or terminal, cyme, generally umbel- or raceme-like, or flowers 1–2. Flower: bisexual, radial; perianth parts, especially petals, overlapped, twisted to right or left, at least in bud; sepals generally 5, fused at base, often reflexed, persistent; petals generally 5, fused in basal ± 1/2; stamens generally 5, attached to corolla tube or throat, alternate lobes, free or fused to form filament column and anther head, filament column then generally with 5 free or fused, ± elaborate appendages abaxially, pollen ± free or removed in pairs of pollinia; nectaries 0 or near ovaries, then 2 or 5, or in stigmatic chambers; ovaries 2, superior or ± so, free [fused]; style tips, stigmas generally fused into massive pistil head. Fruit: 1–2 follicles, (capsule), [berry, drupe]. Seed: many, often with tuft of hairs at 1 or both ends.
200–450 genera, 3000–5000 species: all continents, especially tropics, subtropical South America, southern Africa; many ornamental (including Asclepias, Hoya, Nerium, Plumeria, Stapelia); cardiac glycosides, produced by some members formerly treated in Asclepiadaceae, used as arrow poisons, in medicine to control heart function, and by various insects for defense. [Fishbein 2001 Ann Missouri Bot Gard 88:603–623] Asclepiadaceae ("asclepiads"), although monophyletic, included in Apocynaceae because otherwise the latter is paraphyletic. Complexity of floral structure, variation in asclepiads arguably greatest among all angiosperms. Pattern of carpel fusion (carpels free in ovule-bearing region, fused above), present ± throughout Apocynaceae (in broad sense), nearly unknown in other angiosperms. Base chromosome number generally 11; abundance of latex, generally small size of chromosomes evidently have impeded cytological investigations. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Civeyrel et al. 1998 Molec Phylogen Evol 9:517–527; Rosatti 1989 J Arnold Arbor 70:307–401]
Key to Apocynaceae
1 sp.: California. (Greek: ring gland, from nectary) [Sipes & Wolf 1997 Amer J Bot 84:401–409]
Perennial herb, ± erect, 6–12 cm, fleshy (including large root), herbage tomentose to generally glabrous, glaucous. Leaf: opposite, 2–5 pairs, < 9 cm; petiole < to > blade; blade ovate or ± round, base ± truncate to tapered. Inflorescence: cyme, 2–6-flowered. Flower: > 15 mm; calyx lobes narrow-triangular; corolla 15–20 mm, funnel-shaped, with 5 ± round appendages behind anthers, rose-purple, lobes obovate or round, margins wavy; filaments free, appearing to be attached at base of corolla tube but fused to it up to level of stigma, unappendaged, hairy, anthers forming cone around but free from stigma, each partly sterile, sharply sagittate, pollen ± free; nectaries 5, fused into a 5-lobed disk around but not exceeding ovaries; style thread-like; stigma skirted at base. Fruit: 3–5 cm. Seed: with tuft of long hairs at 1 end.Key to Cycladenia humilis
2n=14. Varieties possibly untenable, merit study. [Online Interchange]
Unabridged note: Given that hairiness within Cycladenia humilis var. humilis appears to be governed by a single gene (thus reducing Cycladenia humilis var. tomentosa to synonymy under Cycladenia humilis var. humilis), doubt is cast on the hair characters separating Cycladenia humilis var. humilis from Cycladenia humilis var. venusta and Cycladenia humilis var. jonesii as well; given that corolla lobe length in Cycladenia humilis var. humilis overlaps that in Cycladenia humilis var. venusta and Cycladenia humilis var. jonesii, doubt is cast on the characters separating these varieties as well. Therefore, the entire group merits further study.
Plant generally glabrous, glaucous, less often dense-tomentose. Flower: perianth glabrous (tomentose) abaxially; corolla glabrous or papillate adaxially, lobes (5)7–9(11) mm.
Loose gravel or sand, talus slopes, often with Pinus ponderosa; 1200–2800 m. Klamath Ranges, High North Coast Ranges, High Cascade Range, n High Sierra Nevada (Plumas, Sierra cos.), Outer South Coast Ranges (Santa Lucia Range). Apr–Aug [Online Interchange]
Unabridged note: Tomentose (assignable to Cycladenia humilis var. tomentosa) and glabrous plants intermixed within populations without intergrading and without showing other differences, suggesting hairiness is governed by a single gene (e.g., JEPS28583, from Lassen Volcanic National Park, with "single gene difference?", in reference to hairiness, on collection label). venusta.
Previous taxon: Cycladenia humilis
Next taxon: Cycladenia humilis var. jonesii
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 28 2015
Citation for this treatment: [Author of taxon treatment] 2013. Cycladenia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=57255, accessed on Aug 28 2015
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|Cycladenia humilis var. humilis|
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© 2007 Aaron Schusteff
|Geographic subdivisions indicated for the distribution of Cycladenia humilis var. humilis|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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