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CONVOLVULACEAE MORNING-GLORY FAMILY

Robert E. Preston, except as noted

Annual, perennial herb, subshrub, generally twining or trailing. Leaf: 0 or alternate. Inflorescence: cyme or flowers 1 in axils; bracts subtending flowers 0 or 2. Flower: bisexual, radial; sepals (4)5, ± free, overlapping, persistent, often unequal; corolla generally showy, generally bell-shaped, ± shallowly 5-lobed, generally pleated and twisted in bud; stamens 5, epipetalous; pistil 1, ovary superior, chambers generally 2, each generally 2-ovuled, styles 1–2. Fruit: generally capsule. Seed: 1–4(6).
55–60 genera, 1600–1700 species: warm temperate to tropics; some cultivated for food or as ornamental (Ipomoea). [Stefanovic et al. 2003 Syst Bot 28:791–806] Monophyletic only if Cuscutaceae included, as treated here. Ipomoea cairica (L.) Sweet, Ipomoea hederacea Jacq. [Ipomoea nil L., misappl.], Ipomoea indica (Burm.) Merr. (including Ipomoea mutabilis Ker Gawl.), Ipomoea purpurea (L.) Roth, Ipomoea triloba L., all included in TJM (1993), not naturalized. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Stefanovic et al. 2002 Amer J Bot 89:1510–1522]

Key to Convolvulaceae

CUSCUTA DODDER

Mihai Costea & Saša Stefanović

Vine, annual (per if on perennial host), not in contact with ground, attached to, holoparasitic on host by many small, specialized roots (haustoria) along stem, generally glabrous. Stem: thread-like, ± green, yellow, orange, or ± red. Leaf: 0 or scale-like, alternate, ± 2 mm. Inflorescence: generally cyme, head- to panicle-like (flowers 1), subtended by 0–3 bracts. Flower: bisexual, radial, parts generally in 4s or 5s; calyx generally divided 2/5–3/5, persistent generally ± cream-white; corolla generally ± white, persistent (withered in fruit) or not, tube cup-shaped to cylindric, bulged or horizontally ridged below lobes or generally not, generally with scales subtending stamens, lobes alternate stamens, erect to reflexed; ovary superior, chambers 2, each 2-ovuled, styles 2, generally free, persistent, stigmas 2, generally spheric, persistent. Fruit: capsule, generally indehiscent to irregularly dehiscent (or circumscissile near base), spheric to ovoid, depressed or not, thickened and/or raised around generally inconspicuous opening between styles or not. Seed: 1–4; coat papillate when hydrated, honeycombed when dry, (rarely neither, with cells ± rectangular, in ± jigsaw-puzzle-like arrangement); embryo generally slender, 1–3-coiled.
± 180 species: cosmopolitan, especially warmer regions of western hemisphere and Polynesia. (Aramaic, Hebrew; to cover, from habit) [Costea & Stefanovic 2009 Syst Bot 34:570–579] By persistent, withered corolla, fruit may be "capped" (corolla on top of fruit), "surrounded" (fruit at least in part visible, corolla ± loosely around fruit), or "enclosed" (fruit not visible, corolla ± tightly around fruit). Cuscuta pentagona Engelm. excluded.
Unabridged etymology: (Aramaic, Hebrew; from the verb K-S-Y (Kaph, Shin, Yodh), to cover, from habit)
Unabridged references: [Costea et al. 2005 Brittonia 57:264–272; Costea et al. 2006 Sida 22:151–175, 177–195, 197–207, 209–225; Costea & Stefanovic 2009. Cuscuta jepsonii (Convolvulaceae), an invasive weed or an extinct endemic? Amer J Bot 96:1744–1750; Costea et al. 2009. Untangling the systematics of salt marsh dodders: Cuscuta pacifica a new segregate species from Cuscuta salina (Convolvulaceae). Syst Bot 34:787–795; Costea & Stefanovic. 2009. Molecular phylogeny of Cuscuta californica complex (Convolvulaceae) and a new species from New Mexico and Trans-Pecos. Syst Bot 34:570–579; Costea & Tardif 2006 Canad J Plant Sci 86:293–316]

Key to Cuscuta

C. obtusiflora Kunth var. glandulosa Engelm.
NATIVE
Inflorescence: spike- or panicle-like, flowers 5–18; pedicels 0–1 mm. Flower: 1.8–2.5 mm, membranous, not papillate, parts in (4s)5s; calyx generally = corolla tube, shallowly cup-shaped, divided 1/2, not veined, not shiny, lobes ovate, bases ± overlapped, margins entire, tip obtuse; corolla tube 1–1.5 mm, bell-shaped, scales reaching stamen bases, oblong, rounded, fringed laterally, densely in apical 2/3, lobes erect to spreading, generally = tube, ovate to ovate-oblong, margins entire, tip obtuse, straight; filaments 0.4–0.6 mm, anthers exserted, 0.3–0.4 mm, ovate to widely elliptic; styles 0.4–1.1 mm, <= ovary. Fruit: 1.5–3 mm, 2.5–4 mm wide, depressed-spheric, not thickened or raised around opening between styles, not translucent, surrounded by corolla. Seed: generally 4, 1.4–1.6 mm, 1.2–1.35 mm wide, widely ovate to widely elliptic.
On herbs including Alternanthera, Dalea, Lythrum, Polygonum, Xanthium; ± < 500 m. Extirpated, formerly sporadically collected in San Bernardino Co., 1890–1898, Sonoma, Butte, Merced cos., 1940s; eastern and southern United States, West Indies, Mexico. Jul–Oct [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 23 2014
Citation for this treatment: [Author of taxon treatment] 2012. Cuscuta, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=57201, accessed on Apr 23 2014

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Bioregions in which Cuscuta obtusiflora var. glandulosa occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.