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Annual to shrub, some aquatic. Leaf: basal or cauline, alternate or opposite (whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0. Inflorescence: raceme, spike, or flowers axillary in 1–few-flowered clusters; flowers few to many, each subtended by 1 bract. Flower: unisexual or bisexual, radial or bilateral; sepals 4–5, generally fused at base; corolla 4–5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1–2, anthers opening by 2 slits; ovary superior, 2–4-chambered, style 1, stigma lobes 0 or 2. Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
110 genera, ± 2000 species: worldwide, especially temperate. [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2001 Molec Phylogen Evol 16:96–112] Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic California genera of Scrophulariaceae (except Buddleja, Limosella, Mimulus, Myoporum, Scrophularia, Verbascum). California Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Key to Plantaginaceae
Annual, often glandular, sometimes brown-staining. Leaf: opposite; proximal petioled. Inflorescence: bracted, often interrupted; flowers 1–many in leaf axils. Flower: calyx lobes 5, generally glabrous on inner surface; corolla ± pea-like, uniformly pale, or generally with pale regions, especially throat and base of upper lip (± uniformly dark in Collinsia greenei), generally glabrous outside, tube short, throat ± angled to tube, ± pouched on upper side, lips generally ± = throat, upper lobes 2, ± reflexed, lower lobes 3, lateral spreading, central lobe keeled, enclosing stamens and style; stamens 4, attached unequally near throat base, spur at base of upper filaments > 1 mm, vestigial, or 0; staminode gland-like; style > 2 mm, stigma minutely 2-lobed. Fruit: septicidal and loculicidal (valves 2-lobed). Seed: generally few, ± oblong, generally plump; inner surface ± concave.Key to Collinsia
± 20 species: North America, especially California. (Zaccheus Collins, Philadelphia botanist, 1764–1831) Late-season flowers generally atypically small.
Plant 5–25 cm. Inflorescence: open, conspicuously densely glandular, staining; distal-most bracts 0–2 mm; pedicels generally >> calyx, some strongly reflexed in fruit, S-shaped. Flower: calyx ± = fruit, lobe tips ± blunt; corolla 4–10 mm, throat angled to tube, as wide as long, throat and lower lip generally blue (lavender ± purple), upper lip ± white or pale lavender, with projections bulging < 1 mm away from throat opening at base of each lobe; filaments glabrous, upper pair sometimes hairy at base, basal spur 0. Seed: 2. Collinsia torreyi var. wrightii and Collinsia torreyi var. brevicarinata perhaps best treated as Collinsia wrightii S. Watson. [Online Interchange]
Leaf: generally 2–5 × longer than wide, elliptic to ovate. Flower: corolla generally 6–9 mm, upper lip with projections bulging ± 0.5 mm away from throat opening at base of each lobe.
Conifer forest, often in sandy soil; 1000–2500 m. Klamath Ranges, High North Coast Ranges, High Cascade Range, Modoc Plateau; southern Oregon. Jun–Aug [Online Interchange]
Previous taxon: Collinsia torreyi var. brevicarinata
Next taxon: Collinsia torreyi var. torreyi
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Mar 9 2014
Citation for this treatment: [Author of taxon treatment] 2012. Collinsia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=56724, accessed on Mar 9 2014
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