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MONTIACEAE MINER'S LETTUCE FAMILY

John M. Miller, except as noted

Annual to perennial herb; generally fleshy. Stem: 1–many, generally glabrous. Leaf: simple, alternate or opposite. Inflorescence: axillary or terminal; cyme, raceme, panicle, umbel, or flower 1. Flower: bisexual, radial; sepals generally 2(9), free; petals (1)2–19, free or ± fused; stamens 1–many, epipetalous or not, anthers pink, rose, or yellow; ovary superior, chamber 1, ovules 1–many, placenta basal or free-central; styles (0)1–8, generally fused at base, branched. Fruit: capsule, circumscissile or 2–3-valved. Seed: 1–many, shiny or ± pebbly or sculptured, black or gray, generally with oil-filled appendage as food for ants.
± 22 genera, ± 230 species: generally temperate America, Asia, Australia, Europe, Kerguelen Is, New Zealand, southern Africa, poorly represented in Europe; some cultivated (Lewisia, Calandrinia). [Ogburn & Edwards 2009 Amer J Bot 96:391–408] Details of flowers, seeds require 20× magnification. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Applequist et al. 2006 Syst Bot 31:310–319; Nyffeler et al. 2008 Haseltonia 14: 26–36]
Unabridged note: May include Hectorellaceae.

Key to Montiaceae

CALYPTRIDIUM PUSSYPAWS

John M. Miller & C. Matt Guilliams

Annual, perennial herb, ± fleshy, from taproot or fibrous roots, glabrous. Stem: 1–several, generally spreading to ascending. Leaf: basal or basal and cauline, simple, oblanceolate to spoon-shaped; basal rosetted. Inflorescence: raceme, panicle, or umbel, scapose, bracts generally < sepals, leaf-like or not; flowers generally on 1 side of axis, persistent in fruit or not; pedicels ± jointed with a transverse groove or constriction. Flower: sepals 2, ovate to reniform, generally scarious or scarious-margined, persistent in fruit; petals 2–4, < sepals, tips adherent, forming cap in fruit (fruit cap), falling as 1 unit; stamens 1–3, anthers pink, rose, or yellow; style included to exserted, stigmas 2. Fruit: 2-valved, generally compressed, narrowly oblong to ± round, generally translucent, deciduous or not. Seed: 1–many, black, dull, fine- to coarse-papillate to shiny, smooth.
8 species: western North America. (Greek: cap, for petal tips in fruit) [Hershkovitz 2006 Gayana Bot 63:13–74]
Unabridged references: [Hershkovitz 1990 Phytologia 68:267–270; Hershkovitz 1991a Ann Missouri Bot Gard 78:1009–1021; Hershkovitz 1991b Phytologia 70:209–225; Hershkovitz 1992 Syst Bot 17:220–238; Hershkovitz & Zimmer 2000 Molec Phylogen Evol 15:419–439; Hinton 1975 Brittonia 27:197–208; Howell 1945 Leafl W Bot 4:214–216; Kelley et al. 2003 FNANM: 4:460–464; Thomas 1956 Leafl W Bot 8:9–11]

Key to Calyptridium

C. parryi A. Gray
NATIVE
Annual 2–11 cm; taproot slender. Stem: spreading to ascending, leafy. Leaf: basal and cauline, 1–3 cm, basal generally withering in fruit. Inflorescence: axillary, raceme or panicle, open to dense, 1–3.5 cm; bracts ovate to elliptic; pedicel stout, <= 1 mm, generally jointed at base; flowers persistent in fruit to not. Flower: sepals 2–5 mm, generally unequal (outer >, wider than inner), round, ovate, or reniform, generally scarious on margin or becoming scarious throughout; petals 4, 1.5–3 mm, generally white; stamens (1)3, anthers pink; style 0.5–1 mm, stigma ± sessile. Fruit: 3–7 mm, ovate to oblong, deciduous or not. Seed: 5–20, 0.5–0.8 mm.
2n=44. [Cistanthe parryi (A. Gray) Hershk.] Unnamed var. in Baja California. [Online Interchange]

C. parryi var. hesseae J.H. Thomas SANTA CRUZ MOUNTAINS PUSSYPAWS
NATIVE
Flower: sepals ovate, margin generally narrowly scarious. Fruit: readily deciduous. Seed: 0.5–0.6 mm, papillae marginal.
Sandy soils in chaparral, oak woodland, conifer forest; 600–1050 m. sw San Francisco Bay Area. [Cistanthe parryi var. hesseae (J.H. Thomas) Kartesz & Gandhi] Apr–Jul [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 24 2014
Citation for this treatment: [Author of taxon treatment] 2013. Calyptridium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=55522, accessed on Oct 24 2014

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Geographic subdivisions indicated for the distribution of Calyptridium parryi var. hesseae Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.