Directory       News       Site Map       Home
    Jepson eFlora: Taxon page
Key to families | Table of families and genera

Previous taxon Indexes to all accepted names and synonyms:
| A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |
Previous taxon


Alan R. Smith, except as noted

Perennial herb in soil or rock crevices; rhizome generally short-creeping, ascending, or erect, scales small to large, generally tan to brown, generally uniformly colored. Leaf: generally tufted or short-spaced, 5–200+ cm, generally ± alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade generally 1–3-pinnate, ± glabrous or with hairs, hair-like scales, or gland-tipped hairs on axes, veins generally free (or netted); rachis, costa generally grooved adaxially. Sporangia: sori round, oblong, J-shaped, or linear along veins; indusia 0 or oblong, J-shaped, reniform, or linear, or of many segmented hair- or scale-like fragments or lobes encircling sorus from below; spores elliptic, winged, ridged, or spiny, scar linear.
± 15 genera, 700 species: worldwide, especially tropics, wooded areas, but some genera (e.g., Cystopteris, Woodsia) generally temperate. See note, reference (Smith et al. 2006 Taxon 55:705–731) under Dryopteridaceae for removal of genera from that family to this. —Scientific Editor: Thomas J. Rosatti.

Key to Woodsiaceae

Rhizome short-creeping to suberect, stout. Leaf: stipe stout, fleshy, easily crushed, straw-colored except base generally blackened, base scaly, ×-section with 2 crescent-shaped vascular strands; blade generally >= 2-pinnate, pinnae of equal sides, ± glabrous or minutely hairy, veins free. Sporangia: sori ± round, ± oblong, or J-shaped; indusia 0, oblong, J-shaped, or reniform, laterally attached.
± 100 species: generally northern temperate, especially eastern Asia. (Greek: doorless, from enclosed sori)

Key to Athyrium

A. filix-femina (L.) Roth var. cyclosorum Rupr.
Leaf: blade elliptic to lanceolate, 1–2-pinnate-pinnatifid, lower 2–4 pinna pairs generally << those above, ultimate segments pinnately lobed to ± toothed, midribs below near base often with minute, branched hairs 0.1–0.2 mm.
2n=80. Woodland, along streams, seepage areas; < 3200 m. California Floristic Province (except Tehachapi Mountain Area, Sacramento Valley, Inner South Coast Ranges, South Coast, Western Transverse Ranges, Peninsular Ranges except San Jacinto Mountains), Modoc Plateau; to Alaska, western Canada, Idaho, Colorado, northern Central America. [Athyrium filix-femina var. californicum Butters; Athyrium filix-femina var. sitchense Rupr.] Highly variable, but named varieties in western North America seem indistinct; other varieties worldwide. [Online Interchange]

Previous taxon: Athyrium distentifolium var. americanum
Next taxon: Cystopteris


Name search

Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 30 2015
Citation for this treatment: [Author of taxon treatment] 2013. Athyrium, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 30 2015

Copyright © 2014 Regents of the University of California
We encourage links to these pages, but the content may not be downloaded for reposting, repackaging, redistributing, or sale in any form, without written permission from The Jepson Herbarium.

click for enlargement Athyrium filix-femina var. cyclosorum
See CalPhotos for additional images
2008 Keir Morse

Geographic subdivisions indicated for the distribution of Athyrium filix-femina var. cyclosorum Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

View elevation by latitude chart
Data provided by the participants of the Consortium of California Herbaria.
View all CCH records


CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.