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Perennial herb in soil or rock crevices; rhizome generally short-creeping, ascending, or erect, scales small to large, generally tan to brown, generally uniformly colored. Leaf: generally tufted or short-spaced, 5–200+ cm, generally ± alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade generally 1–3-pinnate, ± glabrous or with hairs, hair-like scales, or gland-tipped hairs on axes, veins generally free (or netted); rachis, costa generally grooved adaxially. Sporangia: sori round, oblong, J-shaped, or linear along veins; indusia 0 or oblong, J-shaped, reniform, or linear, or of many segmented hair- or scale-like fragments or lobes encircling sorus from below; spores elliptic, winged, ridged, or spiny, scar linear.
± 15 genera, 700 species: worldwide, especially tropics, wooded areas, but some genera (e.g., Cystopteris, Woodsia) generally temperate. See note, reference (Smith et al. 2006 Taxon 55:705–731) under Dryopteridaceae for removal of genera from that family to this. —Scientific Editor: Thomas J. Rosatti.
Key to Woodsiaceae
Rhizome short-creeping to suberect, stout. Leaf: stipe stout, fleshy, easily crushed, straw-colored except base generally blackened, base scaly, ×-section with 2 crescent-shaped vascular strands; blade generally >= 2-pinnate, pinnae of equal sides, ± glabrous or minutely hairy, veins free. Sporangia: sori ± round, ± oblong, or J-shaped; indusia 0, oblong, J-shaped, or reniform, laterally attached.Key to Athyrium
± 100 species: generally northern temperate, especially eastern Asia. (Greek: doorless, from enclosed sori)
Leaf: blade elliptic to lanceolate, 1–2-pinnate-pinnatifid, lower 2–4 pinna pairs generally << those above, ultimate segments pinnately lobed to ± toothed, midribs below near base often with minute, branched hairs 0.1–0.2 mm.
2n=80. Woodland, along streams, seepage areas; < 3200 m. California Floristic Province (except Tehachapi Mountain Area, Sacramento Valley, Inner South Coast Ranges, South Coast, Western Transverse Ranges, Peninsular Ranges), Modoc Plateau; to Alaska, western Canada, Idaho, Colorado, northern Central America. [Athyrium filix-femina var. californicum Butters] Highly variable, but named varieties in western North America seem indistinct; other varieties worldwide. [Online Interchange]
Previous taxon: Athyrium distentifolium var. americanum
Next taxon: Cystopteris
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Mar 7 2014
Citation for this treatment: [Author of taxon treatment] 2012. Athyrium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=54990, accessed on Mar 7 2014
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|Athyrium filix-femina var. cyclosorum|
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Charles Webber © 1999 California Academy of Sciences
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