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WOODSIACEAE CLIFF FERN FAMILY

Alan R. Smith, except as noted

Perennial herb in soil or rock crevices; rhizome generally short-creeping, ascending, or erect, scales small to large, generally tan to brown, generally uniformly colored. Leaf: generally tufted or short-spaced, 5–200+ cm, generally ± alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade generally 1–3-pinnate, ± glabrous or with hairs, hair-like scales, or gland-tipped hairs on axes, veins generally free (or netted); rachis, costa generally grooved adaxially. Sporangia: sori round, oblong, J-shaped, or linear along veins; indusia 0 or oblong, J-shaped, reniform, or linear, or of many segmented hair- or scale-like fragments or lobes encircling sorus from below; spores elliptic, winged, ridged, or spiny, scar linear.
± 15 genera, 700 species: worldwide, especially tropics, wooded areas, but some genera (e.g., Cystopteris, Woodsia) generally temperate. See note, reference (Smith et al. 2006 Taxon 55:705–731) under Dryopteridaceae for removal of genera from that family to this. —Scientific Editor: Thomas J. Rosatti.

Key to Woodsiaceae

ATHYRIUM LADY FERN
Rhizome short-creeping to suberect, stout. Leaf: stipe stout, fleshy, easily crushed, straw-colored except base generally blackened, base scaly, ×-section with 2 crescent-shaped vascular strands; blade generally >= 2-pinnate, pinnae of equal sides, ± glabrous or minutely hairy, veins free. Sporangia: sori ± round, ± oblong, or J-shaped; indusia 0, oblong, J-shaped, or reniform, laterally attached.
± 100 species: generally northern temperate, especially eastern Asia. (Greek: doorless, from enclosed sori)

Key to Athyrium

A. distentifolium Opiz var. americanum (Butters) Cronquist
NATIVE
Leaf: blade narrow-elliptic or lanceolate, 2–3-pinnate, lower 1 or 2 pinna pairs generally ± < those above, ultimate segments shallowly to deeply pinnately lobed, midribs below glabrous or with scattered scales.
2n=80. Moist or wet rock crevices, talus, cliffs, boulder bases, streamsides; 1700–3700 m. Klamath Ranges, High Cascade Range, High Sierra Nevada, Warner Mountains, White and Inyo Mountains; to Alaska, western Canada, Montana, Colorado; also eastern Can, Greenland, eastern Asia. [Athyrium alpestre Milde var. americanum Butters, illeg.] Athyrium distentifolium var. distentifolium in Eurasia. [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 28 2014
Citation for this treatment: [Author of taxon treatment] 2013. Athyrium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=54985, accessed on Nov 28 2014

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Geographic subdivisions indicated for the distribution of Athyrium distentifolium var. americanum Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.