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FABACEAE (Leguminosae) LEGUME FAMILY

Martin F. Wojciechowski, except as noted

Annual to tree. Leaf: generally alternate, generally compound, generally stipuled, generally entire, pinnately veined Inflorescence: generally raceme, spike, umbel or head; or flowers 1–few in axils. Flower: generally bisexual, generally bilateral; hypanthium 0 or flat to tubular; sepals generally 5, generally fused; petals generally 5, free, fused, or lower 2 ± united into keel (see 3, Key to Groups, for banner, wings); stamens 10 or many (or [1], 5, 6, 7, 9), free or fused or 10 with 9 filaments at least partly fused, 1 (uppermost) free; pistil 1, ovary superior, generally 1-chambered, ovules 1–many, style, stigma 1. Fruit: legume, including a stalk-like base (above receptacle) or not. Seed: 1–many, often ± reniform, generally hard, smooth.
± 730 genera, 19400 species: worldwide; with grasses, requisite in agriculture, most natural ecosystems. Many cultivated, most importantly Arachis, peanut; Glycine, soybean; Phaseolus, beans; Medicago, alfalfa; Trifolium, clovers; many orns. [Lewis et al. (eds) 2005 Legumes of the World. RBG, Kew] Unless stated otherwise, fruit length including stalk-like base, number of 2° leaflets is per 1° leaflet. Upper suture of fruit adaxial, lower abaxial. Anthyllis vulneraria L. evidently a waif, a contaminant of legume seed from Europe. Laburnum anagyroides Medik., collected on Mount St. Helena in 1987, may be naturalized. Ceratonia siliqua L., carob tree (Group 2), differs from Gleditsia triacanthos L. in having evergreen (vs deciduous) leaves that are 1-pinnate (vs 1-pinnate on spurs on old stems, 2-pinnate on new stems) with 2–5(8) (vs 7–17) 1° leaflets, commonly cultivated, now naturalized in southern California. Aeschynomene rudis Benth. <Noxious weed>, Halimodendron halodendron (Pall.) Voss <Noxious weed> (possibly extirpated), Lens culinaris Medik. are agricultural weeds. Caragana arborescens Lam. only cultivated. Ononis alopecuroides L. <Noxious weed>, Sphaerophysa salsula (Pall.) DC. <Noxious weed> all evidently extirpated. Cercidium moved to Parkinsonia; Chamaecytisus to Cytisus; Psoralidium lanceolatum to Ladeania. —Scientific Editors: Martin F. Wojciechowski, Thomas J. Rosatti.

Key to Fabaceae

ASTRAGALUS LOCOWEED, MILKVETCH

Martin F. Wojciechowski & Richard Spellenberg

Annual, perennial herb from crown, generally unarmed; hairs generally present, simple or branches 2, from base, parallel to leaf surface, unequal or not. Stem: 0 or prostrate to erect. Leaf: odd-1-pinnate (or palmately compound); leaflets generally jointed to midrib, entire; stipules membranous, lower fused around stem into sheaths (stipule sheaths) or not. Inflorescence: raceme, head- or umbel-like or not, axillary; flowers 2–many. Flower: bilateral; keel petals with small protrusion at base locking into pit on adjacent wing; 9 filaments fused, 1 free; ovary (and fruit) generally sessile, style slender, stigma minute. Fruit: generally 1- or ± 2-chambered, often mottled, generally ± dry in age, sometimes deciduous (falling from plant with or without pedicel, calyx, receptacle) before dehiscence. Seed: 2–many, smooth, compressed, ± notched at attachment scar.
> 2500 species: ± worldwide (380 in North America, 97 in California, including many rare taxa). (Greek: ankle-bone or dice, perhaps from rattling of seeds within fruit) Difficult; flower and fruit needed for identification; fruit said to be "deciduous" dehisce only after fruit has separated from plant; many good species appear similar; some species complexes need study. Taxa near province boundaries may appear in > 1 key. Varieties keyed under species for simplicity; species with varieties so identified in key. Fruit length including beak and any stalk-like base unless fruit body specified; fruit depth is suture-to-suture axis. Astragalus tephrodes A. Gray var. brachylobus (A. Gray) Barneby in southwestern Utah, Arizona, near California.
Unabridged references: [Barneby 1964 Mem New York Bot Gard 20:1–1188; Isely 1998 Native and Naturalized Leguminosae (Fabaceae) of the United States]

Key to Astragalus

A. douglasii (Torr. & A. Gray) A. Gray DOUGLAS MILKVETCH
NATIVE
Perennial herb, leafy; ± minutely strigose. Stem: generally many, ± decumbent or erect, 2–10 dm. Leaf: 5–18 cm; leaflets 7–25, 5–25 mm, elliptic, ovate, or obovate, midrib often raised abaxially, margins ± green, tips obtuse or shallowly notched. Inflorescence: flowers 10–30, spaced, spreading or ascending. Flower: calyx green, lobes 0.7–2.6 mm; petals ± white to pale yellow, banner 8–13 mm, recurved 60–90°, keel 6.2–10.8 mm, keel, wing claws 2.2–4.9 mm, keel blade 4.3–6.4 mm, > keel claw. Fruit: spreading or ± ascending, 25–60 mm, 12–32 mm wide, bladdery, sparsely hairy to glabrous, drying thinly papery; beak erect, pointed, compressed side-to-side; chamber 1; deciduous. Much like Astragalus macrodon (hairs ± spreading, ± wavy; leaflet margins purple); like Astragalus pomonensis (leaflets 25–41; keel blade <= keel claw); varieties ± intergrade. [Online Interchange]

Key to Astragalus douglasii

A. douglasii var. parishii (A. Gray) M.E. Jones
NATIVE
Stem: prostrate to ± ascending, 2–6 dm. Leaf: leaflets 15–25. Flower: calyx tube most densely hairy between lobes, lobes generally 0.7–2.2 mm, triangular, ± as wide as long. Fruit: 25–50 mm.
2n=22. Open areas; 100–2350 m. South Coast, San Bernardino Mountains, Peninsular Ranges. May–Aug [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 24 2014
Citation for this treatment: [Author of taxon treatment] 2012. Astragalus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=54721, accessed on Apr 24 2014

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Bioregions in which Astragalus douglasii var. parishii occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.