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FABACEAE (Leguminosae) LEGUME FAMILY

Martin F. Wojciechowski, except as noted

Annual to tree. Leaf: generally alternate, generally compound, generally stipuled, generally entire, pinnately veined Inflorescence: generally raceme, spike, umbel or head; or flowers 1–few in axils. Flower: generally bisexual, generally bilateral; hypanthium 0 or flat to tubular; sepals generally 5, generally fused; petals generally 5, free, fused, or lower 2 ± united into keel (see 3, Key to Groups, for banner, wings); stamens 10 or many (or [1], 5, 6, 7, 9), free or fused or 10 with 9 filaments at least partly fused, 1 (uppermost) free; pistil 1, ovary superior, generally 1-chambered, ovules 1–many, style, stigma 1. Fruit: legume, including a stalk-like base (above receptacle) or not. Seed: 1–many, often ± reniform, generally hard, smooth.
± 730 genera, 19400 species: worldwide; with grasses, requisite in agriculture, most natural ecosystems. Many cultivated, most importantly Arachis, peanut; Glycine, soybean; Phaseolus, beans; Medicago, alfalfa; Trifolium, clovers; many orns. [Lewis et al. (eds) 2005 Legumes of the World. RBG, Kew] Unless stated otherwise, fruit length including stalk-like base, number of 2° leaflets is per 1° leaflet. Upper suture of fruit adaxial, lower abaxial. Anthyllis vulneraria L. evidently a waif, a contaminant of legume seed from Europe. Laburnum anagyroides Medik., collected on Mount St. Helena in 1987, may be naturalized. Ceratonia siliqua L., carob tree (Group 2), differs from Gleditsia triacanthos L. in having evergreen (vs deciduous) leaves that are 1-pinnate (vs 1-pinnate on spurs on old stems, 2-pinnate on new stems) with 2–5(8) (vs 7–17) 1° leaflets, commonly cultivated, now naturalized in southern California. Aeschynomene rudis Benth. <Noxious weed>, Halimodendron halodendron (Pall.) Voss <Noxious weed> (possibly extirpated), Lens culinaris Medik. are agricultural weeds. Caragana arborescens Lam. only cultivated. Ononis alopecuroides L. <Noxious weed>, Sphaerophysa salsula (Pall.) DC. <Noxious weed> all evidently extirpated. Cercidium moved to Parkinsonia; Chamaecytisus to Cytisus; Psoralidium lanceolatum to Ladeania. —Scientific Editors: Martin F. Wojciechowski, Thomas J. Rosatti.

Key to Fabaceae

ASTRAGALUS LOCOWEED, MILKVETCH

Martin F. Wojciechowski & Richard Spellenberg

Annual, perennial herb from crown, generally unarmed; hairs generally present, simple or branches 2, from base, parallel to leaf surface, unequal or not. Stem: 0 or prostrate to erect. Leaf: odd-1-pinnate (or palmately compound); leaflets generally jointed to midrib, entire; stipules membranous, lower fused around stem into sheaths (stipule sheaths) or not. Inflorescence: raceme, head- or umbel-like or not, axillary; flowers 2–many. Flower: bilateral; keel petals with small protrusion at base locking into pit on adjacent wing; 9 filaments fused, 1 free; ovary (and fruit) generally sessile, style slender, stigma minute. Fruit: generally 1- or ± 2-chambered, often mottled, generally ± dry in age, sometimes deciduous (falling from plant with or without pedicel, calyx, receptacle) before dehiscence. Seed: 2–many, smooth, compressed, ± notched at attachment scar.
> 2500 species: ± worldwide (380 in North America, 97 in California, including many rare taxa). (Greek: ankle-bone or dice, perhaps from rattling of seeds within fruit) Difficult; flower and fruit needed for identification; fruit said to be "deciduous" dehisce only after fruit has separated from plant; many good species appear similar; some species complexes need study. Taxa near province boundaries may appear in > 1 key. Varieties keyed under species for simplicity; species with varieties so identified in key. Fruit length including beak and any stalk-like base unless fruit body specified; fruit depth is suture-to-suture axis. Astragalus tephrodes A. Gray var. brachylobus (A. Gray) Barneby in southwestern Utah, Arizona, near California.
Unabridged references: [Barneby 1964 Mem New York Bot Gard 20:1–1188; Isely 1998 Native and Naturalized Leguminosae (Fabaceae) of the United States]

Key to Astragalus

A. calycosus S. Watson var. calycosus TORREY'S MILKVETCH
NATIVE
Perennial herb, ± stemless, tufted; silvery-strigose, hairs branched. Leaf: 1–7 cm; leaflets 1–7, generally 3, crowded near leaf tip, 5–19 mm, elliptic to obovate, tips generally obtuse or acute. Inflorescence: flowers 1–8, ascending or spreading. Flower: petals ± white to bright purple, wing tips white, notched, banner 10–13 mm, keel 7.4–9.4. Fruit: ascending, 10–25 mm, 3–4 mm wide, oblong, ± 3- sided (lower narrow, grooved), strigose; chambers ± 2.
2n=22. Rocky areas, sagebrush scrub, pine forest; 1500–3550 m. East of Sierra Nevada, n Desert Mountains; to Idaho, Wyoming. Apr–Jul [Online Interchange]
Unabridged note: Expanded author citation: Astragalus calycosus Torr. ex S. Watson var. calycosus

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Dec 22 2014
Citation for this treatment: [Author of taxon treatment] 2013. Astragalus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=54654, accessed on Dec 22 2014

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Geographic subdivisions indicated for the distribution of Astragalus calycosus var. calycosus Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.