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VIOLACEAE VIOLET FAMILY

R. John Little

Annual, perennial herb, [small shrub, tree, vine], from caudices, taproots, rhizomes, or stolons; hairs 0 or simple. Stem: 0 or prostrate to erect. Leaf: basal, cauline, or both, alternate, [opposite], simple to compound, petioled; stipules generally small; blade linear to round, entire to toothed or lobed. Inflorescence: flowers 1 [raceme], axillary or scapose; peduncle bractlets 2, generally alternate. Flower: bisexual, bilateral [radial]; sepals 5, free, basal lobes present [0], generally not prominent; petals 5, free, lowest often largest, base ± elongated into a spur; stamens 5, alternate petals, filaments short, wide, with large in-pointing hairs, lowest 2 anthers with basal nectaries extending into petal spur; ovary superior, chamber 1, placentas parietal, 3, ovules [1] generally many, style 1, often enlarged distally, stigma often oblique or hooked, hairy or not. Fruit: capsule [berry], 3-valved, loculicidal, explosively dehiscent or not. Seed: generally with outgrowth, attractive to ants.
23 genera, 830 species: worldwide, generally temperate, tropics (especially higher elevations). [Munzinger & Ballard 2003 Syst Bot 28:345–351] Lengths of lowest petal including spur. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Baird, V.B. 1942. Wild violets of North Am. University of California Press, Berkeley. Beattie, A. J. and N. Lyons. 1975. Seed dispersal in Viola (Violaceae):adaptations and strategies. Amer. J. Bot. 62: 714–722. Brainerd, E. 1921. Violets of North Am. Vermont Agric. Exp. Sta. Bull. 224. Brizicky, G.K. 1961. The genera of Violaceae in the southeastern United States. J. Arnold Arbor. 42: 321–333. Clausen, J. 1929. Chromosome number and relationship of some North American species of Viola. Annals of Botany. 43: 741–764. Clausen, J. 1964. Cytotaxonomy and distributional ecology of western North American violets. Madroño 17:173–197. Gershoy, A. 1928. Studies in North American violets. I. General considerations. Vermont Agric. Exp. Sta. Bull. 279.]

VIOLA VIOLET
Leaf: generally deciduous. Flower: sepals ± equal, entire; petals unequal, lowest generally largest, with spur generally < 3 [20] mm, lateral 2 equal, generally spreading, upper 2 equal, erect or reflexed, overlapped or not, lateral 2 generally, others sometimes with beard of variously shaped hairs basally; cleistogamous flowers generally present, petals 0. Fruit: ovoid to oblong, hairy or not. Seed: 8–75.
± 500 species: temperate, worldwide, Hawaii, Andes. (Latin: classical name) Important orns including Viola odorata, Viola tricolor L. (Johnny-jump-up, wild pansy), Viola ×wittrockiana Gams (garden pansy).
Unabridged references: [Baker, M.S. 1949a. Studies in western violets. IV. Leafl. West. Bot. 5: 141–147; Baker, M.S. 1949b. Studies in western violets. VI; Madroño 10: 110–128; Baker, M.S. 1957. Brittonia 9: 217–230; Ballard, H.E. 1992. Systematics of Viola Section Viola in North Am north of Mexico. M.S. thesis, Central Michigan University, Mt. Pleasant, Michigan; Ballard, H.E. 1994. Violets of Michigan. Michigan Botanist 33: 131–199; Davidse, G. 1976. A study of some Intermountain violets (Viola Sect. Chamaemelanium). Madroño 23: 274–283; Fabijan et al. 1987. The taxonomy of the Viola nuttallii complex. Can. J. Bot. 65: 2562–2580; Gil-Ad, N.L. 1995. Systematics and evolution of Viola L. subsection Boreali Americanae (W. Becker) Brizicky. Ph.D. dissertation, Univ of Michigan, Ann Arbor, Michigan; Gil-Ad, N.L. 1997. Systematics of Viola subsection Boreali-Americanae. Boissiera 53: 1–130; Hitchcock, C.L. 1961. Vasc. Plant. Pacific NorthW. Vol. 3; McKinney, L.E. 1992. A taxonomic revision of the acaulescent blue violets (Viola) of North Am. Sida Botanical Miscellany, 7: 1–59; McKinney, L.E. and N. Russell. 2002. Violaceae of the Southeastern United States. Castanea 4: 369–379; Russell, N.H. 1965. Violets (Viola) of the central and eastern United States: an introductory survey. Sida 1: 1–113]

Key to Viola

V. purpurea Kellogg
NATIVE
Perennial herb. Stem: prostrate to erect, generally several, clustered on 1–several subterranean caudices from woody rhizome. Leaf: simple, base tapered or truncate to cordate, often oblique; basal 1–6 per caudex, stipule oblong, fused to, forming 2 membranous wings on petiole, entire or few-toothed, the tip of each wing free, deltate or lanceolate, generally fringed, blade 0.8–5.3 cm, 0.4–4.1 cm wide, lanceolate to round; cauline petiole 0.3–12.3 cm, blade 0.3–4.8 cm, 0.3–5 cm wide, ovate, oblong, elliptic, triangular, diamond-shaped or lanceolate, crenate, serrate or dentate with 3–4(5) prominent pointed or rounded lobes per side, shallowly dentate, sharp angled, or entire, occasionally wavy. Inflorescence: axillary; peduncle 1–17 cm. Flower: sepals lanceolate, ciliate or not; petals deep lemon-yellow, upper 2, sometimes lateral 2 red- to purple-brown abaxially, lower 3 veined dark brown, lateral 2 bearded with club-shaped hairs, lowest petal 6–16 mm. Fruit: 4–12 mm, ovoid to ± spheric, puberulent. Seed: 2.1–3.1 mm, light to dark brown or gray, mottled brown.
2n=12,24. Subspecies variable, intergrade, need study; no additional subspecies [Online Interchange]
Unabridged note: Subspecies variable, intergrade, need study; no additional subspecies; plants in shade often produce larger leaves, longer stems.

V. purpurea subsp. quercetorum (M.S. Baker & J.C. Clausen) R.J. Little
NATIVE
Plant 4–25(34.5) cm, puberulent. Stem: generally not buried, spreading to erect, generally elongated by end of season. Leaf: basal petiole 1.9–9.5 cm, blade 1–5.3 cm, 1–3.5 cm wide, ovate to round, ± unlobed, irregularly wavy-dentate, generally gray-green, occasionally purple-tinted abaxially, base tapered to truncate or ± cordate, tip obtuse; cauline petiole 1.3–5.3 cm, blade 1.4–2.2 cm, 0.8–1.3 cm wide, lanceolate to diamond-shaped, crenate-serrate, base generally tapered, tip acute. Inflorescence: peduncle 2.7–17 cm. Flower: lowest petal 10–16 mm. Fruit: 8–12 mm.
2n=24. Dry, grassy or brushy slopes, chaparral, generally below yellow-pine forest; 304–1981 m. Northwestern California, Cascade Range, Sierra Nevada, Central Western California, Southwestern California, Modoc Plateau; southwestern Oregon. Fresh material (light gray) ± needed to distinguish from Viola purpurea subsp. purpurea (dull to bright green tinted ± purple); perhaps better treated as a separate sp. Feb–Jul [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 23 2014
Citation for this treatment: [Author of taxon treatment] 2012. Viola, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=53410, accessed on Apr 23 2014

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click for enlargement Viola purpurea subsp. quercetorum
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Beatrice F. Howitt 1999 California Academy of Sciences

Bioregions in which Viola purpurea subsp. quercetorum occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.