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Key to families | Table of families and genera
Indexes to all accepted names and synonyms:
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Annual, perennial herb, [small shrub, tree, vine], from caudices, taproots, rhizomes, or stolons; hairs 0 or simple. Stem: 0 or prostrate to erect. Leaf: basal, cauline, or both, alternate, [opposite], simple to compound, petioled; stipules generally small; blade linear to round, entire to toothed or lobed. Inflorescence: flowers 1 [raceme], axillary or scapose; peduncle bractlets 2, generally alternate. Flower: bisexual, bilateral [radial]; sepals 5, free, basal lobes present , generally not prominent; petals 5, free, lowest often largest, base ± elongated into a spur; stamens 5, alternate petals, filaments short, wide, with large in-pointing hairs, lowest 2 anthers with basal nectaries extending into petal spur; ovary superior, chamber 1, placentas parietal, 3, ovules  generally many, style 1, often enlarged distally, stigma often oblique or hooked, hairy or not. Fruit: capsule [berry], 3-valved, loculicidal, explosively dehiscent or not. Seed: generally with outgrowth, attractive to ants.
23 genera, 830 species: worldwide, generally temperate, tropics (especially higher elevations). [Munzinger & Ballard 2003 Syst Bot 28:345–351] Lengths of lowest petal including spur. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Baird, V.B. 1942. Wild violets of North Am. University of California Press, Berkeley. Beattie, A. J. and N. Lyons. 1975. Seed dispersal in Viola (Violaceae):adaptations and strategies. Amer. J. Bot. 62: 714–722. Brainerd, E. 1921. Violets of North Am. Vermont Agric. Exp. Sta. Bull. 224. Brizicky, G.K. 1961. The genera of Violaceae in the southeastern United States. J. Arnold Arbor. 42: 321–333. Clausen, J. 1929. Chromosome number and relationship of some North American species of Viola. Annals of Botany. 43: 741–764. Clausen, J. 1964. Cytotaxonomy and distributional ecology of western North American violets. Madroño 17:173–197. Gershoy, A. 1928. Studies in North American violets. I. General considerations. Vermont Agric. Exp. Sta. Bull. 279.]
Leaf: generally deciduous. Flower: sepals ± equal, entire; petals unequal, lowest generally largest, with spur generally < 3  mm, lateral 2 equal, generally spreading, upper 2 equal, erect or reflexed, overlapped or not, lateral 2 generally, others sometimes with beard of variously shaped hairs basally; cleistogamous flowers generally present, petals 0. Fruit: ovoid to oblong, hairy or not. Seed: 8–75.Key to Viola
± 500 species: temperate, worldwide, Hawaii, Andes. (Latin: classical name) Important orns including Viola odorata, Viola tricolor L. (Johnny-jump-up, wild pansy), Viola ×wittrockiana Gams (garden pansy).
Unabridged references: [Baker, M.S. 1949a. Studies in western violets. IV. Leafl. West. Bot. 5: 141–147; Baker, M.S. 1949b. Studies in western violets. VI; Madroño 10: 110–128; Baker, M.S. 1957. Brittonia 9: 217–230; Ballard, H.E. 1992. Systematics of Viola Section Viola in North Am north of Mexico. M.S. thesis, Central Michigan University, Mt. Pleasant, Michigan; Ballard, H.E. 1994. Violets of Michigan. Michigan Botanist 33: 131–199; Davidse, G. 1976. A study of some Intermountain violets (Viola Sect. Chamaemelanium). Madroño 23: 274–283; Fabijan et al. 1987. The taxonomy of the Viola nuttallii complex. Can. J. Bot. 65: 2562–2580; Gil-ad, N.L. 1995. Systematics and evolution of Viola L. subsection Boreali-Americanae (W. Becker) Brizicky. Ph.D. dissertation, Univ of Michigan, Ann Arbor, Michigan; Gil-ad, N.L. 1997. Systematics of Viola subsection Boreali-Americanae. Boissiera 53: 1–130; Gil-ad, N.L. 1998. The micromorphologies of seed coats and petal trichomes of the taxa of Viola subsect. Boreali-Americanae (Violaceae) and their utility in discerning orthospecies from hybrids. Brittonia 50: 91–121; Hitchcock, C.L. 1961. Vasc. Plant. Pacific NorthW. Vol. 3; McKinney, L.E. 1992. A taxonomic revision of the acaulescent blue violets (Viola) of North Am. Sida Botanical Miscellany, 7: 1–59; McKinney, L.E. and N. Russell. 2002. Violaceae of the Southeastern United States. Castanea 4: 369–379; Russell, N.H. 1965. Violets (Viola) of the central and eastern United States: an introductory survey. Sida 1: 1–113]
Perennial herb 7.5–30 cm, glabrous to densely puberulent. Stem: prostrate to erect, generally several, clustered on 1–several subterranean caudices from woody rhizome. Leaf: simple; basal 1–5 per caudex, petiole 4.3–19.2 cm, blade 2.3–8.5 cm, 1.4–3.7 cm wide, elliptic or ovate, entire, wavy, or generally irregularly crenate to serrate, base tapered, often oblique to ± truncate, tip acute or obtuse; cauline blade 2.6–5.8 cm, 1.3–3.5 cm wide, like basal. Inflorescence: axillary; peduncle 2.7–26 cm. Flower: sepals lanceolate, ciliate or not; petals deep lemon-yellow, upper 2, sometimes lateral 2 maroon or ± brown abaxially, lower 3 veined brown-purple, lateral 2 bearded with cylindric hairs, lowest 12–20 mm. Fruit: 6–12 mm, elliptic to oblong, glabrous to minutely puberulent. Seed: 2–3 mm, medium to dark brown, ± 1/3 length covered by outgrowth.
2n=36,48. Study needed; 1 other subsp., ± northwestern United States. [Online Interchange]
Unabridged note: Complex; study needed; hybrids obscure subspecies; 1 other subsp., Idaho, Montana, Utah, Washington, Wyoming.
Plant 7.5–21 cm. Leaf: basal and cauline, blade generally ovate, base generally ± truncate; basal 2.3–6.7 cm, not much longer than wide, ± crenate or serrate, wavy, or entire; cauline blade 1.3–3 cm wide. Inflorescence: peduncle 5.5–26 cm. Seed: dark brown.
2n=36. Generally in vernally moist soil, slopes, meadows, conifer forest, sagebrush; 122–2440 m. Northwestern California, Cascade Range, n&c High Sierra Nevada, Warner Mountains; to British Columbia, Alberta, Utah. [Viola praemorsa Douglas ex Lindl. subsp. praemorsa; Viola nuttallii Pursh subsp. praemorsa (Douglas ex Lindl.) Piper; Viola nuttallii Pursh var. praemorsa (Douglas ex Lindl.) S. Watson] Mar–Jul [Online Interchange]
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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 26 2015
Citation for this treatment: [Author of taxon treatment] 2013. Viola, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=53398, accessed on Nov 26 2015
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|Geographic subdivisions indicated for the distribution of Viola praemorsa subsp. praemorsa|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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