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| Jepson eFlora: Taxon page
Key to families | Table of families and genera |
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Indexes to all accepted names and synonyms: | A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z | |
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Annual, perennial herb, [small shrub, tree, vine], from caudices, taproots, rhizomes, or stolons; hairs 0 or simple.
Stem: 0 or prostrate to erect.
Leaf: basal, cauline, or both, alternate, [ opposite], simple to compound, petioled; stipules generally small; blade linear to round, entire to toothed or lobed.
Inflorescence: flowers 1 [ raceme], axillary or scapose; peduncle bractlets 2, generally alternate.
Flower: bisexual, bilateral [ radial]; sepals 5, free, basal lobes present [0], generally not prominent; petals 5, free, lowest often largest, base ± elongated into a spur; stamens 5, alternate petals, filaments short, wide, with large in-pointing hairs, lowest 2 anthers with basal nectaries extending into petal spur; ovary superior, chamber 1, placentas parietal, 3, ovules [1] generally many, style 1, often enlarged distally, stigma often oblique or hooked, hairy or not.
Fruit: capsule [ berry], 3-valved, loculicidal, explosively dehiscent or not.
Seed: generally with outgrowth, attractive to ants.
23 genera, 830 species: worldwide, generally temperate, tropics (especially higher elevations). [Munzinger & Ballard 2003 Syst Bot 28:345–351] Lengths of lowest petal including spur. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Baird, V.B. 1942. Wild violets of North Am. University of California Press, Berkeley. Beattie, A. J. and N. Lyons. 1975. Seed dispersal in Viola (Violaceae):adaptations and strategies. Amer. J. Bot. 62: 714–722. Brainerd, E. 1921. Violets of North Am. Vermont Agric. Exp. Sta. Bull. 224. Brizicky, G.K. 1961. The genera of Violaceae in the southeastern United States. J. Arnold Arbor. 42: 321–333. Clausen, J. 1929. Chromosome number and relationship of some North American species of Viola. Annals of Botany. 43: 741–764. Clausen, J. 1964. Cytotaxonomy and distributional ecology of western North American violets. Madroño 17:173–197. Gershoy, A. 1928. Studies in North American violets. I. General considerations. Vermont Agric. Exp. Sta. Bull. 279.]
Key to Viola
Leaf: generally deciduous.
Flower: sepals ± equal, entire; petals unequal, lowest generally largest, with spur generally < 3 [20] mm, lateral 2 equal, generally spreading, upper 2 equal, erect or reflexed, overlapped or not, lateral 2 generally, others sometimes with beard of variously shaped hairs basally; cleistogamous flowers generally present, petals 0.
Fruit: ovoid to oblong, hairy or not.
Seed: 8–75.
± 500 species: temperate, worldwide, HI, Andes. (Latin: classical name) Important orns including Viola odorata, Viola tricolor L. (Johnny-jump-up, wild pansy), Viola ×wittrockiana Gams (garden pansy).
Unabridged references: [Baker, M.S. 1949a. Studies in western violets. IV. Leafl. West. Bot. 5: 141–147; Baker, M.S. 1949b. Studies in western violets. VI; Madroño 10: 110–128; Baker, M.S. 1957. Brittonia 9: 217–230; Ballard, H.E. 1992. Systematics of Viola Section Viola in North Am north of Mexico. M.S. thesis, Central Michigan University, Mt. Pleasant, Michigan; Ballard, H.E. 1994. Violets of Michigan. Michigan Botanist 33: 131–199; Davidse, G. 1976. A study of some Intermountain violets (Viola Sect. Chamaemelanium). Madroño 23: 274–283; Fabijan et al. 1987. The taxonomy of the Viola nuttallii complex. Can. J. Bot. 65: 2562–2580; Gil-Ad, N.L. 1995. Systematics and evolution of Viola L. subsection Boreali Americanae (W. Becker) Brizicky. Ph.D. dissertation, Univ of Michigan, Ann Arbor, Michigan; Gil-Ad, N.L. 1997. Systematics of Viola subsection Boreali-Americanae. Boissiera 53: 1–130; Hitchcock, C.L. 1961. Vasc. Plant. Pacific NorthW. Vol. 3; McKinney, L.E. 1992. A taxonomic revision of the acaulescent blue violets (Viola) of North Am. Sida Botanical Miscellany, 7: 1–59; McKinney, L.E. and N. Russell. 2002. Violaceae of the Southeastern United States. Castanea 4: 369–379; Russell, N.H. 1965. Violets (Viola) of the central and eastern United States: an introductory survey. Sida 1: 1–113]
Perennial 7.5–30 cm, glabrous to densely puberulent.
Stem: prostrate to erect, generally several, clustered on 1–several subterranean caudices from woody rhizome.
Leaf: simple; basal 1–5 per caudex, petiole 4.3–19.2 cm, blade 2.3–8.5 cm, 1.4–3.7 cm wide, elliptic or ovate, entire, wavy, or generally irregularly crenate to serrate, base tapered, often oblique to ± truncate, tip acute or obtuse; cauline blade 2.6–5.8 cm, 1.3–3.5 cm wide, like basal.
Inflorescence: axillary; peduncle 2.7–26 cm.
Flower: sepals lanceolate, ciliate or not; petals deep lemon-yellow, upper 2, sometimes lateral 2 maroon or ± brown abaxially, lower 3 veined brown-purple, lateral 2 bearded with cylindric hairs, lowest 12–20 mm.
Fruit: 6–12 mm, elliptic to oblong, glabrous to minutely puberulent.
Seed: 2–3 mm, medium to dark brown, ± 1/3 length covered by outgrowth.
2n=36,48. Study needed; 1 other subsp., ± nw US. [Online Interchange]
Unabridged note: Complex; study needed; hybrids obscure subspp.; 1 other subsp., ID, MT, UT, WA, WY.
Plant 7.5–21 cm.
Leaf: basal and cauline, blade generally ovate, base generally ± truncate; basal 2.3–6.7 cm, not much longer than wide, ± crenate or serrate, wavy, or entire; cauline blade 1.3–3 cm wide.
Inflorescence: peduncle 5.5–26 cm.
Seed: dark brown.
2n=36. Generally in vernally moist soil, slopes, meadows, conifer forest, sagebrush; 122–2440 m. Northwestern California, Cascade Range, n&c High Sierra Nevada, Warner Mountains;
Previous taxon: Viola praemorsa subsp. linguifolia
Next taxon: Viola primulifolia subsp. occidentalis
Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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| Bioregions in which taxon occurs | Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues. |
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Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates. | Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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