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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

STREPTANTHUS JEWELFLOWER
Annual to perennial herb, generally ± glaucous; hairs simple or 0. Leaf: basal rosetted or not, petioled, entire or dentate to pinnately lobed or divided; cauline sessile, occasionally petioled, base generally lobed or clasping. Inflorescence: elongated. Flower: radial or bilateral; calyx urn- or occasionally bell-shaped, sepals erect, base ± sac-like, keeled or not; petal blade narrower to wider than proximal 1/2, generally channeled, margins ± crinkled or not; stamens in 3 pairs of unequal length, or 4 long and 2 short, longest filaments fused or free. Fruit: silique, dehiscent, linear, flat parallel to septum, unsegmented; stigma entire or 2-lobed. Seed: 10–120, in 1 row, generally winged.
35 species: southwestern United States, northern Mexico. (Greek: twisted flower, from wavy-margined petals)

Key to Streptanthus

S. insignis Jeps. SAN BENITO JEWELFLOWER
NATIVE
Annual, bristly throughout. Stem: 1.2–6 dm, simple or branched distally. Leaf: basal not rosetted, lanceolate to oblanceolate, coarsely dentate to pinnately lobed; mid-cauline sessile, 1.3–9 cm, lanceolate, dentate, basally lobed; distal reduced, dentate. Inflorescence: open; terminal cluster of dark purple, pale yellow, or ± green-yellow sterile flowers present. Flower: calyx bell-shaped, sepals of fertile flowers 4–6 mm, oblong to lanceolate, not or ± keeled, dark purple, ± purple, or ± green-yellow, hairy; petals 7–12 mm, 1–2 mm wide, ± purple-white, lemon-yellow, or ± yellow-white, midvein darker; filaments in 3 pairs of unequal length; longest pair completely fused, 6–9 mm, lower pair free; fertile anthers 1.3–2 mm. Fruit: ascending or reflexed, 3.5–11.4 cm, 1.5–2 mm wide, straight or ± curved, not constricted between seeds; valves sparsely to densely bristly, midvein prominent; stigma entire; pedicels spreading to ascending, 3–8 mm. Seed: 32–94, 1.4–2 mm, broadly oblong; wing 0.1–0.3 mm wide, continuous.
2n=28. [Online Interchange]

S. insignis subsp. lyonii Kruckeb. & J.L. Morrison ARBURUA RANCH JEWELFLOWER
NATIVE
Inflorescence: terminal sterile flower cluster pale yellow, ± green-yellow, or ± purple. Flower: sepals of fertile flowers ± green-yellow or ± purple; petals lemon yellow or ± yellow-white. Fruit: moderately to densely bristly-hairy.
Serpentine, grassland, oak woodland; 200–900 m. Inner South Coast Ranges (near Ortigalita Peak, w Merced Co.). Apr–May [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 20 2014
Citation for this treatment: [Author of taxon treatment] 2013. Streptanthus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=53146, accessed on Sep 20 2014

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Geographic subdivisions indicated for the distribution of Streptanthus insignis subsp. lyonii Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.