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Robert Patterson, family description, key to genera

Annual, perennial herb, shrub, vine. Leaf: simple or compound, cauline (or most basal), alternate or opposite; stipules 0. Inflorescence: cymes, heads, clusters, or flower 1; bracts in involucres or not. Flower: sepals generally 5, fused at base, translucent membrane generally connecting lobes, torn by fruit; corolla generally 5-lobed, radial or bilateral, salverform to bell-shaped, throat often well defined; stamens generally 5, epipetalous, attached at >= 1 level, filaments of >= 1 length, pollen white, yellow, blue, or red; ovary superior, chambers generally 3, style 1, stigmas generally 3. Fruit: capsule. Seed: 1–many, when wetted swelling or not, gelatinous or not.
26 genera, 314 species: America, northern Europe, northern Asia; some cultivated (Cantua, Cobaea (cup-and-saucer vine), Collomia, Gilia, Ipomopsis, Linanthus, Phlox). [Porter & Johnson 2000 Aliso 19:55–91] Leptodactylon moved to Linanthus. —Scientific Editors: Robert Patterson, Thomas J. Rosatti.

Key to Polemoniaceae


Leigh A. Johnson

Annual, generally prickly. Stem: prostrate to generally erect; branches ascending or spreading; glabrous to generally hairy, often glandular. Leaf: simple, alternate (lowermost opposite), entire to generally deeply pinnate-lobed. Inflorescence: generally head-like, bracts pinnate- to palmate-toothed or -lobed, spine-tipped (flower 1–2, pedicels elongate, bracts entire, not spine-tipped). Flower: calyx lobes 4–5, equal, entire or toothed, or unequal, spine-tipped; corolla lobes 4–5; stigmas 2 or 3. Fruit: generally ovoid, chambers 1–3; dehiscing when wetted, seeds adherent to fruit and each other, or generally dehiscing at maturity, seeds free. Seed: 1–many per fruit, brown, gelatinous when wet.
± 40 species: western North America, Argentina, Chile. (F.F. Navarrete, Spanish physician, ?–1742) [Porter & Johnson 2000 Aliso 19:55–91] Revised taxonomy, too late for full treatment in TJM2, includes Navarretia linearifolia (Howell) L.A. Johnson subsp. linearifolia, a ± cryptic segregate of Navarretia sinistra, and Navarretia linearifolia subsp. pinnatisecta (H. Mason & A.D. Grant) L.A. Johnson [Navarretia sinistra subsp. pinnatisecta] (Johnson & Cairns-Heath 2010 Syst Bot 35:618–628); Navarretia paradoxinota and Navarretia paradoxiclara, both new to science, and Navarretia propinqua [Navarretia intertexta subsp. propinqua] (Johnson et al. 2013 Phytotaxa 91:27–38). Relative positions of flower parts are as pressed, unless stated otherwise.
Unabridged references: [Porter 1996 Aliso 15:57–77; Spencer & Porter 1997 Syst Bot 22:649–668]

Key to Navarretia

N. hamata Greene
Plant erect, 1° axes generally 1–5; odor generally skunk-like. Stem: 8–30 cm, branches ascending; glandular-puberulent. Leaf: pinnate-lobed; axis linear to widely lanceolate; lobes spreading; tip resembling a grappling hook, with 3 divergent spiny lobes. Inflorescence: bracts widely clasping, outer lanceolate, recurved, pinnate-lobed, generally with 3 divergent spiny lobes at tip ± separated from proximal lobes. Flower: calyx lobes entire or toothed; corolla generally purple or ± pink; stigmas 3. Fruit: < calyx, dehiscing tip to base. Subspecies intergrade. [Online Interchange]

N. hamata subsp. hamata
Leaf: cauline axis linear or lanceolate. Inflorescence: generally terminal; bracts gland-dotted or glandular-hairy, lobes generally unequal. Flower: calyx lobes entire or toothed; corolla tube included, lobes 3–5 mm.
2n=18. Dry, sandy, rocky places in coastal, inland chaparral; < 1200 m. Transverse Ranges, Peninsular Ranges. Plants in Transverse Ranges with wider leaves, axillary inflorescence much like Navarretia hamata subsp. parviloba but with larger, brighter flowers. Apr–Jun [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 26 2015
Citation for this treatment: [Author of taxon treatment] 2013. Navarretia, Revision 1, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 26 2015

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click for enlargement Navarretia hamata subsp. hamata
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2007 Aaron Schusteff

Geographic subdivisions indicated for the distribution of Navarretia hamata subsp. hamata Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.