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LAMIACEAE (Labiatae) MINT FAMILY

Dieter H. Wilken & Margriet Wetherwax, family description, key to genera

Annual to shrub [tree, vine], glabrous to hairy, generally aromatic. Stem: generally erect, generally 4-angled. Leaf: generally simple to deeply lobed, generally opposite, generally gland-dotted. Inflorescence: generally cymes, generally many in dense axillary clusters surrounding stem, generally separated by evident internodes or collectively crowded, spike- or panicle-like, occasionally head-like or raceme, subtended by leaves or bracts; flowers sessile or pedicelled. Flower: generally bisexual; calyx generally 5-lobed, radial to bilateral; corolla generally bilateral, 1–2-lipped, upper lip entire or 2-lobed, ± flat to hood-like, occasionally 0, lower lip generally 3-lobed; stamens generally 4, epipetalous, generally exserted, paired, pairs generally unequal, occasionally 2, staminodes 2 or 0; ovary superior, generally 4-lobed to base chambers 2, ovules 2 per chamber, style 1, generally arising from center at junction of lobes, stigmas generally 2. Fruit: generally 4 nutlets, generally ovoid to oblong, smooth.
± 230 genera, 7200 species: worldwide. Many cultivated for herbs, oils (Lavandula, lavender; Mentha, mint; Rosmarinus, rosemary; Thymus, thyme), some cultivated as ornamental (in California Cedronella, Leonotis, Monarda, Phlomis). [Harley et al. 2004 Fam Generally Vasc Plant 7:167–275] Moluccella laevis L., shell flower, historical waif in California. Satureja calamintha (L.) Scheele subsp. ascendens (Jordan) Briq. reported as alien but not naturalized. Salazaria moved to Scutellaria; California Hyptis moved to Condea, California Satureja moved to Clinopodium. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Lamiaceae

MONARDELLA

Andrew C. Sanders, Mark A. Elvin & Mark S. Brunell

Annual to shrub, ± gland-dotted, scented. Leaf: entire to serrate, margin flat or wavy; petioles 0 or present, often grading into blade. Inflorescence: flowers in compact clusters of >= 1 per main stem, these occasionally arrayed in panicles (rarely spikes); flowers 3–100 per cluster; bracts generally erect in a cup-like involucre or reflexed, reduced in size inward, leaf-like to membranous in texture, green or straw-colored to rose or purple, linear to ovate, acuminate to acute or obtuse. Flower: calyx 5-lobed; 4–25 mm; corolla white to purple or yellow to red, weakly bilateral, upper lip erect, 2-lobed, lower lip recurved, 3-lobed; stamens 4; style unequally 2-lobed.
> 30 species: western North America. (Latin: small Monarda) [Elvin & Sanders 2009 Novon 19:315–343; Epling 1925 Ann Missouri Bot Gard 12:1–106; Jepson 1943 Fl California 431–444] Complex; study needed; many taxa intergrade; flower cluster width and bract orientation given for pressed specimens. Leaf length including petiole, if present. M. Brunell authored Monardella follettii (in part), Monardella odoratissima (in part), Monardella palmeri, Monardella purpurea, Monardella sheltonii (in part), Monardella stebbinsii, Monardella villosa (in part).
Unabridged references: [Abrams 1951 Ill Fl Pac Sts V III:648–660]

Key to Monardella

M. villosa Benth. COYOTE-MINT
NATIVE
Subshrub, matted to erect, open, rhizomed; hairs generally > 0.5 mm, appressed to spreading, sparse to dense, soft, wavy, or woolly, glandular and not, branched or not. Leaf: 10–30 mm (including petiole, 5–10 mm), length < 1–3.3 × width, lanceolate to ovate or widely triangular-ovate, serrate or crenate (entire), base truncate to obtuse or tapered; abaxially glandular-hairy, woolly or not. Inflorescence: flower clusters generally 1–6(10) per main stem, 10–40 mm wide; bracts 10–30 mm, reflexed; bracts generally leaf-like in texture, color, hairiness (or innermost of middle series ± scarious proximally), innermost linear to elliptic-ovate. Flower: calyx tube hairy, glandular, hairs on lobes generally similar to hairs on tube (or stiff-spreading); corolla 10–20 mm, white or pink to purple, lobes obtuse. [Online Interchange]
Unabridged note: Leaf length-to-width ratio > 3 in inland northern Outer North Coast Ranges (Humboldt Co.), <= 1 in southern North Coast and Central Coast. Flower clusters occasionally 6–10 per main stem in Klamath Ranges, northern North Coast Ranges. If recognized taxonomically, plants of coastal foothills of Santa Lucia Range, from San Luis Obispo Co. northern and possibly to Santa Cruz Co., with purple scarious bracts, and ovate, glabrous to sparsely hairy leaves assignable to Monardella villosa var. subglabra Hoover. Literature reports of hybridization involving that entity, Monardella palmeri, and Monardella villosa subsp. obispoensis in the Alder Creek area of Monterey Co. have not been confirmed.

M. villosa subsp. obispoensis (Jeps.) Jokerst
NATIVE
Plant ascending to erect, 15–30 cm, tufted to matted in age; hairs simple and branched, branched hairs many on stem, leaves. Leaf: 15–25 mm, 8–15 mm wide, ± narrowly ovate, hairy, base tapered to obtuse; adaxially green; abaxially white-tomentose, often obscuring leaf surface. Inflorescence: flower cluster 1, 15–25 mm wide, generally subtended by unmodified leaves; bracts ovate. Flower: corolla white to lavender, tube exserted.
Chaparral, oak woodland; < 1500 m. s Central Coast, s Outer South Coast Ranges. Intergrades with Monardella hypoleuca subsp. hypoleuca. May–Aug [Online Interchange]
Unabridged synonyms: [Monardella villosa var. obispoensis Hoover ex Jeps.]
Unabridged note: Intergrades with Monardella palmeri in the vicinity of Cuesta Pass, San Luis Obispo Co. Branched hairs are also known from Monardella villosa in the vicinity of Hoopa Valley, Humboldt Co., vicinity of Dos Rios, Mendocino Co., and vicinity of Idria, San Benito Co., and in Monardella palmeri (bract marginal hairs). Branched hairs also occur in Monardella viridis in Sonoma Co., and occasionally in intergrades with Monardella hypoleuca in Santa Barbara Co. Expanded author citation: Monardella villosa subsp. obispoensis (Hoover ex Jeps.) Jokerst

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 2 2014
Citation for this treatment: [Author of taxon treatment] 2013. Monardella, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=51628, accessed on Sep 2 2014

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Geographic subdivisions indicated for the distribution of Monardella villosa subsp. obispoensis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.