|
|||||||||
Apocynaceae Habit: Annual, perennial herb, shrub, tree, often vine; sap generally milky. Leaf: simple, alternate, opposite, subwhorled to whorled, entire; stipules 0 or small, finger-like. Inflorescence: axillary or terminal, cyme, generally umbel- or raceme-like, or flowers 1--2. Flower: bisexual, radial; perianth parts, especially petals, overlapped, twisted to right or left, at least in bud; sepals generally 5, fused at base, often reflexed, persistent; petals generally 5, fused in basal +- 1/2; stamens generally 5, attached to corolla tube or throat, alternate lobes, free or fused to form filament column and anther head, filament column then generally with 5 free or fused, +- elaborate appendages abaxially, pollen +- free or removed in pairs of pollinia; nectaries 0 or near ovaries, then 2 or 5[10], or in stigmatic chambers; ovaries 2, superior or +- so, free [fused]; style tips, stigmas generally fused into massive pistil head. Fruit: 1--2 follicles, (capsule), [berry, drupe]. Seed: many, often with tuft of hairs at 1 or both ends. Genera In Family: 200--450 genera, 3000--5000 species: all continents, especially tropics, subtropical South America, southern Africa; many ornamental (including Asclepias, Hoya, Nerium, Plumeria, Stapelia); cardiac glycosides, produced by some members formerly treated in Asclepiadaceae, used as arrow poisons, in medicine to control heart function, and by various insects for defense. Note: Asclepiadaceae ("asclepiads"), although monophyletic, included in Apocynaceae because otherwise the latter is paraphyletic. Complexity of floral structure, variation in asclepiads arguably greatest among all angiosperms. Pattern of carpel fusion (carpels free in ovule-bearing region, fused above), present +- throughout Apocynaceae (in broad sense), nearly unknown in other angiosperms. Base chromosome number generally 11; abundance of latex, generally small size of chromosomes evidently have impeded cytological investigations. Jepson eFlora Author: Thomas J. Rosatti Reference: Fishbein 2001 Ann Missouri Bot Gard 88:603--623 Unabridged Reference: Civeyrel et al. 1998 Molec Phylogen Evol 9:517--527; Rosatti 1989 J Arnold Arbor 70:307--401 Scientific Editor: Bruce G. Baldwin. Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Key to Apocynaceae Previous taxon: Yabea microcarpa Next taxon: Amsonia |