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Key to families | Table of families and genera

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POACEAE (Gramineae) GRASS FAMILY

James P. Smith, Jr., except as noted

Annual to woody per; roots generally fibrous.
Stem: generally round, hollow; nodes swollen, solid.
Leaf: alternate, 2-ranked, generally linear, parallel-veined; sheath generally open; ligule membranous or hairy, at blade base.
Inflorescence: various (of generally many spikelets).
Spikelet: glumes generally 2; florets ( lemma, palea, flower) 1–many; lemma generally membranous, sometimes glume-like; palea generally ± transparent, ± enclosed by lemma.
Flower: generally bisexual, minute; perianth vestigial; stamens generally 3; stigmas generally 2, generally plumose.
Fruit: grain (rarely achene-like).
650–900 genera; ± 10550 species: worldwide; greatest economic importance of any family (wheat, rice, maize, millet, sorghum, sugar cane, forage crops, ornamental, weeds; thatching, weaving, building materials). [Barkworth et al. 2003 FNANM:25; Barkworth et al. 2007 FNANM:24] Generally wind-pollinated. Achnatherum, Ampelodesmos, Hesperostipa, Nassella, Piptatherum, Piptochaetium, Ptilagrostis moved to Stipa; Elytrigia, Leymus, Pascopyrum, Pseudoroegneria, Taeniatherum to Elymus; Hierochloe to Anthoxanthum; Lolium, Vulpia to Festuca; Lycurus to Muhlenbergia; Monanthochloe to Distichlis; Pleuraphis to Hilaria; Rhynchelytrum to Melinis. The following taxa (in genera not included here), recorded in CA from historical collections or reported in literature, are extirpated, lacking vouchers, or not considered naturalized: Acrachne racemosa (Roth) Ohwi, Allolepis texana (Vasey) Soderstr. & H.F. Decker, Amphibromus nervosus (Hook. f.) Baill., Axonopus affinis Chase, Axonopus fissifolius (Raddi) Kuhlm., Coix lacryma-jobi L., Cutandia memphitica (Spreng.) K. Richt., Dinebra retroflexa (Vahl) Panz., Eremochloa ciliaris (L.) Merr., Eustachys distichophylla (Lag.) Nees, Gaudinia fragilis (L.) P. Beauv., Miscanthus sinensis Andersson, Neyraudia arundinacea (L.) Henrard, Phyllostachys aurea Rivière & C. Rivière, Phyllostachys bambusoides Siebold & Zuccarini, Rottboellia cochinchinensis (Lour.) Clayton, Schedonnardus paniculatus (Nutt.) Branner & Coville, Schizachyrium cirratum (Hack.) Wooton & Standl., Schizachyrium scoparium (Michx.) Nash, Themeda quadrivalvis (L.) Kuntze, Thysanolaena latifolia (Hornem.) Honda, Tribolium obliterum (Hemsl.) Renvoize, Zea mays L., Zizania palustris L. var. interior (Fassett) Dore, Zoysia japonica Steud. Paspalum pubiflorum E. Fourn., Paspalum quadrifarium Lam., are now reported for s CA (J Bot Res Inst Texas 4:761–770). See Glossary p. 30 for illustrations of general family characteristics. —Scientific Editors: James P. Smith, Jr., J. Travis Columbus, Dieter H. Wilken.
Unabridged references: [Hitchcock 1951 Manual grasses US, USDA Misc Publ 200; Clayton & Renvoise 1986 Kew Bull Add Series 13]

Key to Poaceae

AGROPYRON CRESTED WHEAT GRASS
Perennial, generally cespitose.
Stem: erect or bent, 2.5–10 dm.
Leaf: sheath open, generally appendaged; ligule membranous; blade flat or rolled.
Inflorescence: spike-like, axis not breaking apart at maturity; spikelets 1, 2-ranked, strongly overlapping, divergent or spreading.
Spikelet: laterally compressed, glumes ± equal, < floret, lanceolate, 1–5-veined, keeled, acute to short-awned; florets 3–8[16]; axis breaking above glumes and between florets; lemma 5–7-veined, keeled, acute to awned; palea ± = lemma; anthers 3, 3–5 mm.
12–15 species: Medit, e Eur, c Asia. (Greek: field wheat, perhaps referring to a weed resembling wheat) [Barkworth 2007 FNANM 24:277–279] Siberian wheat grass, Agropyron fragile (Roth) P. Candargy, has also been reported for CA, but is doubtfully naturalized; often used for soil stabilization on range and cropland.
Unabridged references: [Barkworth & Dewey 1985 Amer J Bot 72:767–776]

A. cristatum (L.) Gaertn. subsp. pectinatum (M. Bieb.) Tzvelev
NATURALIZED
Occ rhizomatous.
Stem: generally erect.
Leaf: blade 3–12(20) cm, 1.5–6 mm wide, glabrous or pubescent.
Inflorescence: 1.3–10 cm; internodes generally 1–5 mm, equal or not, glabrous or long-hairy; spikelets diverging at 30–95° angles.
Spikelet: 7–16 mm; glumes 3–6 mm, generally 3-veined, generally awned, awns 1.5–3 mm; florets 3–8; lemma 5–9 mm, generally 5-veined, tip acute, generally awned, awn 1–6 mm.
2n=14,28,42. Disturbed areas, degraded agricultural sites; 600–1500 m. Klamath Ranges, Cascade Range Foothills, n High Sierra Nevada, s Outer South Coast Ranges, Southwestern California, Great Basin Floristic Province, Mojave Desert; most of North America; native to Europe, Mediterranean, Asia. [Agropyron cristatum subsp. desertorum (Link) Á. Löve; Agropyron desertorum (Link) Schult.] Used to rejuvenate burned or overgrazed areas. Jun–Aug [Online Interchange]
Unabridged synonyms: [Agropyron cristatum subsp. desertorum (Fisch. ex Link) Á. Löve; Agropyron desertorum (Fisch. ex Link) Schult.]
Unabridged note: There are Consortium records that, if verified, would voucher elevations up to 2533 m. The following (and possibly other) accessions, if verified, would represent range extensions (as indicated): UCD51678 (NCoRI); SJSU7566 (c SNH); UCD51676 (s SNH).

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.