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Key to families | Table of families and genera
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Perennial herb in soil or rock crevices; rhizome generally short-creeping, ascending, or erect, scales small to large, generally tan to brown, generally uniformly colored. Leaf: generally tufted or short-spaced, 5–200+ cm, generally ± alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade generally 1–3-pinnate, ± glabrous or with hairs, hair-like scales, or gland-tipped hairs on axes, veins generally free (or netted); rachis, costa generally grooved adaxially. Sporangia: sori round, oblong, J-shaped, or linear along veins; indusia 0 or oblong, J-shaped, reniform, or linear, or of many segmented hair- or scale-like fragments or lobes encircling sorus from below; spores elliptic, winged, ridged, or spiny, scar linear.
± 15 genera, 700 species: worldwide, especially tropics, wooded areas, but some genera (e.g., Cystopteris, Woodsia) generally temperate. See note, reference (Smith et al. 2006 Taxon 55:705–731) under Dryopteridaceae for removal of genera from that family to this. —Scientific Editor: Thomas J. Rosatti.
Key to Woodsiaceae
Rhizome generally ascending to suberect, short, old stipe bases many. Leaf: often glandular or hairy; stipe base ×-section with 2 vascular strands; blade 1–2-pinnate, segments ± toothed to pinnately lobed, veins free, ending just short of margin. Sporangia: sori round, generally not at margins; indusium cup-like, often of many segmented hair- or scale-like fragments or lobes encircling sorus from below, often of crusty, ± white beads, often obscure in age.Key to Woodsia
± 30 species: generally northern temperate. (J. Woods, Britain, b. 1776) [Windham 1993 FNANM 2:270–280]
Unabridged references: [Brown 1964 Beih Nova Hedwigia 16:1–154]
Leaf: 5–25 cm, 1–3.5 cm wide, tip ± acute, unforked; hairs on abaxial leaf axes 0 or ± 0.1 mm, cylindric, non-segmented, glandular; pinnae 0.5–2.5 cm, 0.3–1.3 cm wide, pinnately lobed to 1-pinnate, margin fine-toothed. Sporangia: indusium of segmented hairs.
2n=76,152. Crevices, rock bases; 900–2800 m. Klamath Ranges, High Cascade Range, n&s High Sierra Nevada, San Bernardino Mountains, Peninsular Ranges, Modoc Plateau, White and Inyo Mountains, Desert Mountains; to British Columbia, eastern Canada, northern United States, Oklahoma, Arizona. Variable. Sterile, nonglandular plants distinguished from Cystopteris fragilis by leaf veins that do not quite reach the margin in Woodsia but do in Cystopteris. If recognized taxonomically, plants differing in chromosome number, spore size [Windham 1993], perhaps other ways, assignable to W. oregana subsp. cathcartiana (B.L. Rob.) Windham (2n = 152), W. oregana subsp. oregana (2n = 76); study needed. [Online Interchange]
Unabridged note: Variable. Like Cystopteris fragilis except glandular hairs generally present in Woodsia, absent in Cystopteris; veins ending just before leaf margin in Woodsia at 10×, at leaf margin in Cystopteris.
Previous taxon: Woodsia
Next taxon: Woodsia plummerae
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Mar 5 2015
Citation for this treatment: [Author of taxon treatment] 2013. Woodsia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=48612, accessed on Mar 5 2015
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© 2006 Steve Matson
|Geographic subdivisions indicated for the distribution of Woodsia oregana|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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