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VIOLACEAE VIOLET FAMILY

R. John Little

Annual, perennial herb, [small shrub, tree, vine], from caudices, taproots, rhizomes, or stolons; hairs 0 or simple. Stem: 0 or prostrate to erect. Leaf: basal, cauline, or both, alternate, [opposite], simple to compound, petioled; stipules generally small; blade linear to round, entire to toothed or lobed. Inflorescence: flowers 1 [raceme], axillary or scapose; peduncle bractlets 2, generally alternate. Flower: bisexual, bilateral [radial]; sepals 5, free, basal lobes present [0], generally not prominent; petals 5, free, lowest often largest, base ± elongated into a spur; stamens 5, alternate petals, filaments short, wide, with large in-pointing hairs, lowest 2 anthers with basal nectaries extending into petal spur; ovary superior, chamber 1, placentas parietal, 3, ovules [1] generally many, style 1, often enlarged distally, stigma often oblique or hooked, hairy or not. Fruit: capsule [berry], 3-valved, loculicidal, explosively dehiscent or not. Seed: generally with outgrowth, attractive to ants.
23 genera, 830 species: worldwide, generally temperate, tropics (especially higher elevations). [Munzinger & Ballard 2003 Syst Bot 28:345–351] Lengths of lowest petal including spur. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Baird, V.B. 1942. Wild violets of North Am. University of California Press, Berkeley. Beattie, A. J. and N. Lyons. 1975. Seed dispersal in Viola (Violaceae):adaptations and strategies. Amer. J. Bot. 62: 714–722. Brainerd, E. 1921. Violets of North Am. Vermont Agric. Exp. Sta. Bull. 224. Brizicky, G.K. 1961. The genera of Violaceae in the southeastern United States. J. Arnold Arbor. 42: 321–333. Clausen, J. 1929. Chromosome number and relationship of some North American species of Viola. Annals of Botany. 43: 741–764. Clausen, J. 1964. Cytotaxonomy and distributional ecology of western North American violets. Madroño 17:173–197. Gershoy, A. 1928. Studies in North American violets. I. General considerations. Vermont Agric. Exp. Sta. Bull. 279.]

VIOLA VIOLET
Leaf: generally deciduous. Flower: sepals ± equal, entire; petals unequal, lowest generally largest, with spur generally < 3 [20] mm, lateral 2 equal, generally spreading, upper 2 equal, erect or reflexed, overlapped or not, lateral 2 generally, others sometimes with beard of variously shaped hairs basally; cleistogamous flowers generally present, petals 0. Fruit: ovoid to oblong, hairy or not. Seed: 8–75.
± 500 species: temperate, worldwide, Hawaii, Andes. (Latin: classical name) Important orns including Viola odorata, Viola tricolor L. (Johnny-jump-up, wild pansy), Viola ×wittrockiana Gams (garden pansy).
Unabridged references: [Baker, M.S. 1949a. Studies in western violets. IV. Leafl. West. Bot. 5: 141–147; Baker, M.S. 1949b. Studies in western violets. VI; Madroño 10: 110–128; Baker, M.S. 1957. Brittonia 9: 217–230; Ballard, H.E. 1992. Systematics of Viola Section Viola in North Am north of Mexico. M.S. thesis, Central Michigan University, Mt. Pleasant, Michigan; Ballard, H.E. 1994. Violets of Michigan. Michigan Botanist 33: 131–199; Davidse, G. 1976. A study of some Intermountain violets (Viola Sect. Chamaemelanium). Madroño 23: 274–283; Fabijan et al. 1987. The taxonomy of the Viola nuttallii complex. Can. J. Bot. 65: 2562–2580; Gil-Ad, N.L. 1995. Systematics and evolution of Viola L. subsection Boreali Americanae (W. Becker) Brizicky. Ph.D. dissertation, Univ of Michigan, Ann Arbor, Michigan; Gil-Ad, N.L. 1997. Systematics of Viola subsection Boreali-Americanae. Boissiera 53: 1–130; Hitchcock, C.L. 1961. Vasc. Plant. Pacific NorthW. Vol. 3; McKinney, L.E. 1992. A taxonomic revision of the acaulescent blue violets (Viola) of North Am. Sida Botanical Miscellany, 7: 1–59; McKinney, L.E. and N. Russell. 2002. Violaceae of the Southeastern United States. Castanea 4: 369–379; Russell, N.H. 1965. Violets (Viola) of the central and eastern United States: an introductory survey. Sida 1: 1–113]

Key to Viola

V. cuneata S. Watson NORTHERN TWO-EYED VIOLET, WEDGE-LEAVED VIOLET
NATIVE
Perennial herb 2–25 cm, glabrous. Stem: prostrate to erect, generally several from thin, shallow rhizome or a deep caudex with fleshy roots. Leaf: simple; basal 2–6 per caudex, petiole 4.5–9.8 cm, blade 1–4 cm, 2.7–3.8 cm wide, round-ovate to deltate, generally shiny, ± leathery, veined purple, ± serrate, base wedge-shaped, tip acute; cauline petiole 0.5–20 cm, blade 0.9–2.6 cm, 0.7–1.8 cm wide, round to generally diamond-shaped, crenate to ± serrate, base wedge-shaped, tip acute to obtuse. Inflorescence: axillary; peduncle 1–10.5 cm. Flower: sepals lanceolate with purple midvein, not ciliate; petals white adaxially, deep red-violet abaxially, upper 2 often overlapped, with yellow area basally, lateral 2 with large purple spot basally above smaller yellow area, bearded with club-shaped hairs, lowest 8–14 mm, veined purple, with large yellow spot basally. Fruit: 5–9 mm, ± round, glabrous to ± minutely scabrous. Seed: 2.5–3 mm, deep brown-purple.
Uncommon. Vernally moist areas in open pine or oak forests, often on serpentine; 120–2200 m. Northwestern California, n High Sierra Nevada; southwestern Oregon. Mar–Sep [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 16 2014
Citation for this treatment: [Author of taxon treatment] 2013. Viola, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=48200, accessed on Sep 16 2014

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click for enlargement Viola cuneata
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2006 Steve Matson

Geographic subdivisions indicated for the distribution of Viola cuneata Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.