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LENTIBULARIACEAE BLADDERWORT FAMILY

Barry A. Rice

Annual, perennial herb, carnivorous, of moist or aquatic habitats. Stem: caudex or stolon, then often with thread-like branches. Leaf: simple, in rosette, or simple or dissected, emerging from caudex or stolon, with minute, carnivorous bladders ± throughout. Inflorescence: raceme or 1-flowered, scapose. Flower: bisexual; calyx lips 2[4], upper 3-lobed, lower 2-lobed, or lips unlobed; corolla 2-lipped, spurred at base, lower lip flat or arched upward, blocking throat or not; stamens 2, epipetalous; ovary superior, chamber 1, placenta generally free-central; stigma unequally 2-lobed, ± sessile. Fruit: capsule, round, 2-valved, circumscissile, or irregularly dehiscent. Seed: generally many, small.
3 genera, 330 species: worldwide, especially tropics. [Rice 2006 Growing Carnivorous Plants. Timber Press] —Scientific Editor: Thomas J. Rosatti.

Key to Lentibulariaceae

UTRICULARIA BLADDERWORT
Carnivorous by bladders (here treated as modified leaves), into which small organisms are sucked when hairs at opening are triggered [epiphytic]. Stem: submersed or subterranean shoots [rarely caudex]; some aquatic species produce 2 kinds of stems, green (in water or at surface; leaves with flattened or thread-like segments; bladders 0–few) and white (generally buried in substrate; leaves 0; bladders many), the latter not always present in poor collections. Leaf: simple or generally dissected into narrow segments, alternate on stolon, margins often with bristles (visible at 10–30×). Inflorescence: raceme or 1-flowered, emergent; scape slender or wiry, bracts present. Flower: calyx lips 2[4], unlobed; corolla yellow [or not], with red-brown streaks or not, upper lip ± entire, lower entire or 3-lobed, spurred; rarely cleistogamous. Fruit: capsule.
± 220 species: worldwide, especially tropics, Australia. (Latin: little bag, from bladders) [Taylor 1989 Kew Bull Add Ser 14:1–724] Size variable, often unreliable for identification; distinction between stems, leaves uncertain. Glands inside bladders consist of 2 pairs of oppositely directed arms, angles of divergence between which used (at 150× or more) to identify fresh or pressed specimens.

Key to Utricularia

U. intermedia Hayne FLAT-LEAVED BLADDERWORT
NATIVE
Stem: of 2 kinds, some freely floating, green, with leaves, rarely with bladders, others rooted in mud, white, without leaves, with bladders; winter buds bristly. Leaf: 3–15 mm, 3-parted at base, variously dissected above; ultimate segments < 20, ± linear, flattened, margins with 5–12(20) bristles, ± entire, tip obtuse. Inflorescence: 1–5-flowered; peduncle < 25 cm, < 1 mm diam; pedicel erect in fruit. Flower: corolla 10–16 mm; lower lip ± 2 × upper, ± > or ± = cylindric, acute spur. Fruit: circumscissile. Seed: not seen.
2n=44. Shallow (< 1 m) water; 1200–2700 m. Cascade Range, High Sierra Nevada, Modoc Plateau; northern United States, Canada; circumboreal. In some populations, divergence of bladder gland arm-pairs resembling that of Utricularia ochroleuca. Jul–Sep [Online Interchange] {CNPS list}

Previous taxon: Utricularia gibba
Next taxon: Utricularia macrorhiza

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 23 2014
Citation for this treatment: [Author of taxon treatment] 2012. Utricularia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=47604, accessed on Apr 23 2014

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click for enlargement Utricularia intermedia
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© 2009 Barry Rice

Bioregions in which Utricularia intermedia occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.