Jepson Herbarium
		University of California, Berkeley

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Indexes to all accepted names and synonyms: | A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |

Tree.
Leaf: simple, alternate, 2-ranked; veins pinnate; stipules deciduous.
Flower: radial; sepals 4–9, free to fused; corolla 0; stamens 4–9, opposite sepals; ovary superior, chamber 1, ovule 1, style branches 2.
Fruit: 2-winged nutlet.
7 genera, ± 60 species: temperate to tropics; some cultivated for ornamental (Ulmus, Zelkova), used for wood (especially Ulmus). [Sytsma et al. 2002 Ann Missouri Bot Gard 89:1531–1546] Celtis moved to Cannabaceae. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Elias 1970 J Arnold Arbor 51:18–40]

Deciduous.
Leaf: serrate (or doubly so), base generally oblique, 2° veins straight, parallel, extending to margin, each ending in a tooth; axils of 2° veins generally with prominent tufts of hairs.
Inflorescence: umbels or short racemes in leaf axils on old wood; flowers sessile or pedicels 7–17 mm.
Flower: bisexual; calyx generally bell-shaped, lobes 4–9; stamens 4–9, exserted; ovary strongly compressed; style divided to base, branches spreading.
± 40 species: n temperate. Widely cultivated as street trees; flowers, fruit needed for identification.

To 25 m; bark deeply ridged, remaining firmly attached, medium gray.
Stem: corky outgrowths on branches 0; winter buds dark brown or red-brown, ± spheric to ovoid; inner bud scale margins generally white ciliate.
Leaf: 2–8 cm, 1.2–3.5 cm wide, ovate- to lance- elliptic, acute or short- acuminate ( acuminate), margins simply or doubly serrate, abaxial surface glabrous or pubescent only on major veins and tufted in vein axils.
Inflorescence: flower, fruit before leaves in spring; pedicel ± 0.
Fruit: 1–2 cm, 1–1.5 cm wide, ± round (broadly obovate or elliptic), ± white-tan, glabrous except for pubescence on stigmatic surface in notch.
Streambanks, washes, bottomland, roadsides, disturbed areas; 20–1500 m. Sierra Nevada, Southwestern California, Great Basin Floristic Province;

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.