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NYCTAGINACEAE FOUR O'CLOCK FAMILY

Andy Murdock, except as noted

Annual, perennial herb, subshrub, [shrub, tree], glabrous or hairy. Stem: often forked. Leaf: opposite, sessile or petioled, pairs generally unequal; blade generally entire. Inflorescence: generally forked; spike, head-like cluster, or umbel, flowers rarely 1, bracts forming a calyx-like involucre or not. Flower: bisexual, generally ± radial (bilateral), sometimes cleistogamous in some genera; perianth of 1 whorl, generally petal-like, bell- to trumpet-shaped, base hardened, tightly surrounding ovary in fruit, lobes 3–5, generally notched to ± lobed; stamens 1–many; ovary superior (appearing inferior due to hardened perianth base), style 1. Fruit: achene in hardened perianth base; round to ± flat; smooth, angled, ribbed, or winged; glabrous, hairy, or glandular.
30 genera, 350 species: warm regions, especially America; some ornamental (Bougainvillea; Mirabilis, four o'clock). [Spellenberg 2003 FNANM 4:14–17] —Scientific Editor: Thomas J. Rosatti.

Key to Nyctaginaceae

TRIPTEROCALYX
Perennial herb from large taproot. Stem: much-branched. Leaf: petioled; blade < 8 cm, fleshy, margin often wavy. Inflorescence: head or umbel; bracts 5–10, green; flowers maturing 1 side of inflorescence before the other; receptacle flat or rounded, studded with peg-shaped pedicel-like projections 1–3 mm. Flower: nocturnal, perianth trumpet-shaped, lobes 4–5; stamens 3–5, included; stigma linear, included. Fruit: wings 2–4, wide, surrounding fruit body, translucent, conspicuously net-veined.
4 species: arid North America. (Greek, Latin: 3-winged cup, from fruit) [Galloway 2003 FNANM 4:70–71] Closely related to Abronia.

Key to Tripterocalyx

T. micranthus (Torr.) Hook. SMALL-FLOWERED SAND-VERBENA
NATIVE
Stem: < 6 dm, glandular-sticky or scabrous. Leaf: blade 1–6 cm, narrowly ovate to elliptic, glabrous to glandular-hairy, hairs denser abaxially. Flower: perianth generally green-white to light pink.
Sand dunes; 800–2450 m. Mojave Desert; to Montana, South Dakota, New Mexico. Apr–May [Online Interchange] {CNPS list}

Previous taxon: Tripterocalyx crux-maltae
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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 1 2014
Citation for this treatment: [Author of taxon treatment] 2013. Tripterocalyx, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=47318, accessed on Oct 1 2014

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click for enlargement Tripterocalyx micranthus
See CalPhotos for additional images
1979 Dean Wm. Taylor

Geographic subdivisions indicated for the distribution of Tripterocalyx micranthus Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.