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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

THELYPODIUM
Annual to perennial herb; hairs 0 or simple. Leaf: basal rosetted, petioled, entire to pinnately lobed; mid-cauline petioled or sessile, base lobed to sagittate or wedge-shaped. Flower: sepals erect to reflexed, bases sac-like or not; petals linear to oblanceolate, spoon-shaped, or obovate, clawed or not, white to lavender or purple; stamens free (± fused). Fruit: silique, dehiscent, linear, unsegmented, ± narrowed between seeds, cylindric or ± flat parallel to septum, stalked above receptacle; stigma entire. Seed: 1 row per chamber, ± flat; wing generally 0.
16 species: western North America. (Greek: female foot, from fruit stalk above receptacle) [Al-Shehbaz 1973 Contr Gray Herb 204:1–148]

Key to Thelypodium

T. crispum Payson
NATIVE
Biennial, generally glaucous. Stem: 1–7(12) dm; hairs 0 or toward base. Leaf: basal blade 2–15(25) cm, dentate to pinnately lobed (entire), withered by fruit time, petiole ciliate; mid-cauline sessile, base lobed or sagittate, entire or dentate. Inflorescence: dense. Flower: petals 6–11(14.5) mm, 0.5–0.7(1) mm wide, linear to narrowly oblanceolate, white to pale lavender, crinkled. Fruit: 1–2.5(4) cm, 0.7–1(1.8) mm wide, cylindric, narrowed between seeds; stalk above receptacle 0.5–1.5(3.5) mm, slender; style 0.5–1.5(2.5) mm; pedicel erect to erect-ascending, partly or fully appressed, 2–5(10) mm, slender. Seed: 22–50, 1–1.5 mm, plump.
2n=26. Alkaline or sandy soils, lake margins, scrub; 1200–3000 m. High Sierra Nevada, Great Basin Floristic Province; Nevada. Jun–Aug [Online Interchange]
Unabridged note: Expanded author citation: Thelypodium crispum Greene ex Payson

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 29 2014
Citation for this treatment: [Author of taxon treatment] 2013. Thelypodium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=46355, accessed on Jul 29 2014

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click for enlargement Thelypodium crispum
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2007 Steve Matson

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.