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STAPHYLEACEAE BLADDERNUT FAMILY

Robert E. Preston & James R. Shevock

[Perennial herb], shrub, tree; [some monoecious or dioecious]. Leaf: generally opposite, pinnately compound, generally stipuled; leaflets generally toothed. Inflorescence: panicle [raceme], drooping. Flower: radial; sepals 5, free or fused at base, often petal-like; petals 5, free; stamens 5, alternate petals, often attached to disk; ovary superior, chambers 2–4, each with 1–12 ovules in 2 rows. Fruit: generally inflated capsule with open top [follicles, drupe, berry]. Seed: generally 1–2 per chamber.
5 genera, 50–60 species: northern temperate, Asia, Central America, South America; some cultivated for showy fruit (Staphylea), timber (Turpinia). [Matthews & Endress 2005 Bot J Linn Soc 147:1–46] Most closely related to Crossosomataceae, Stachyuraceae. —Scientific Editor: Thomas J. Rosatti.

STAPHYLEA BLADDERNUT
Leaf: deciduous; leaflets generally 3. Flower: petals white or pink. Fruit: bladdery, deeply 3-lobed, pendent. Seed: spheric, light brown.
10 species: northern temperate. (Greek: cluster, from inflorescence)

S. bolanderi A. Gray SIERRA BLADDERNUT
NATIVE
Shrub, small tree, 2–6 m, glabrous. Leaf: leaflets 2.5–6 cm, widely ovate to round, finely serrate. Inflorescence: generally appearing before or with leaves. Flower: sepals 8–10 mm, white; petals 10–12 mm, white; stamens well exserted. Fruit: 2.5–5 cm, prominently horned. Seed: 5–7 mm, ± obovoid, light brown, smooth.
Wooded or shrubby slopes; 240–1720 m. Cascade Range, c&s Sierra Nevada. Mar–May [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Dec 21 2014
Citation for this treatment: [Author of taxon treatment] 2013. Staphylea, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=45426, accessed on Dec 21 2014

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Geographic subdivisions indicated for the distribution of Staphylea bolanderi Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.