Jepson Herbarium
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Perennial [ shrub, tree], [generally glandular-hairy].
Leaf: simple, alternate but generally clustered at stem tips, petioled, stipules 0, margins entire [spiny-toothed].
Inflorescence: generally terminal, [rarely flowers 1 in axils].
Flower: bisexual [ unisexual, plants dioecious], radial, parts generally in 5s, generally with glandular dots, lines, or pits; sepals free or united at base; corolla united at base, lobes spreading; stamens epipetalous, opposite corolla lobes, anthers forming cone as flower opens, then spreading; staminodes 5, alternate stamens, epipetalous, scale-, petal-, or stamen-like; ovary partly inferior, 1-chambered, placenta free-central or basal, style 1, stigma head-like.
Fruit: capsule [generally dry berry or drupe].
Seed: 1–many, generally yellow to orange-red.
7–9 genera, ± 100 species: generally tropical America. [Caris & Smets 2004 Amer J Bot 91:627–643] Incl in Primulaceae in TJM (1993). —Scientific Editor: Thomas J. Rosatti.
Unabridged note: Incl in Primulaceae in TJM (1993). The partially inferior ovary as well as aspects of staminode anatomy are unusual for Primulaceae. Recent research (Källersjö et al. 2000 Amer J Bot 87:1325–1341) has shown that Samolus is a sister-taxon to Theophrastaceae, providing evidence for its removal from Primulaceae.

Erect, glabrous.
Leaf: cauline or basal and cauline, alternate.
Inflorescence: raceme, terminal and axillary.
Flower: corolla > calyx, white.
Fruit: spheric, valves 5, opposite calyx lobes.
± 10 species: ± worldwide. (Celtic: from curative properties)

Stem: 1.5–4 dm.
Leaf: 2–5 cm; blade lance- oblong to obovate, generally tapered to winged petiole, tip obtuse to rounded.
Inflorescence: pedicels ascending to spreading, 1–2 cm, with 1 bractlet ± near middle.
Flower: calyx 1–2 mm, lobes triangular; corolla ± 1.5 mm across.
Fruit: ± 2.5 mm wide.
2n=26. Moist sites; generally < 1300 m. Great Central Valley, Central Western California, South Coast, n Channel Islands, Western Transverse Ranges, San Bernardino Mountains, Peninsular Ranges;

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.