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Shrub, tree; dioecious (monoecious). Stem: trunk < 40 m; wood soft; bark smooth, bitter; buds scaly. Leaf: simple, alternate, deciduous; stipules generally present, deciduous or not, often large. Inflorescence: catkin [or various, or flowers 1]; each flower subtended by 1 bract. Flower: perianth modified into non-nectariferous, cup- or saucer-shaped structure or reduced to adaxial nectary (rarely also with abaxial nectary, then free or fused into shallow cup). Staminate flower: stamens 2–many. Pistillate flower: pistil 1, ovary superior to 1/2-inferior, chambers generally 2–4, placentas parietal, stigma lobes 2–4. Fruit: berry, drupe, or 2–4-valved capsule. Seed: often with basal tuft of hairs.
58 genera, 1210 species: widespread in tropics, northern temperate, arctic. Now including many genera (e.g., Flacourtia, Idesia, Xylosma) formerly in Flacourtiaceae, at least in part because of presence on leaf margins in both families of salicoid teeth (vein extending to tooth tip). In California (and generally outside California), Populus pollinated by wind, Salix by insects, wind. Hybrids common; identification often difficult. —Scientific Editor: Thomas J. Rosatti.
Key to Salicaceae
Shrub, tree; dioecious; bud scale 1, not sticky, margins generally fused (or free, overlapping). Stem: twigs generally flexible, generally not glaucous. Leaf: generally alternate; stipules generally vestigial on first leaves, leaf-like on later; mature blade linear to broadly obovate, entire to toothed, generally ± hairy; petiole glands generally 0. Inflorescence: catkin, blooming before, with, or after leaves, sessile or terminating ± short leafy or bracted shoot ("on leafy shoot" or "on bracted shoot"; "catkin length" including leafless or bractless part of subtending shoot); 1 flower bract subtending each flower, deciduous or persistent, brown, black, or 2-colored (paler proximally, darker distally; darker generally ± brown). Flower: perianth reduced to adaxial nectary (rarely also with abaxial nectary, then free or fused into shallow cup). Staminate flower: stamens (1)2(10); nectary generally 1. Pistillate flower: ovary stalked or sessile, style generally 1, stigmas 2, each 2-lobed, deciduous or persistent; nectary generally 1, generally rod-like. Fruit: valves 2.Key to Salix
± 450 species: ± worldwide, especially northern temperate, arctic. (Latin: ancient name) [Argus 1997 Madroño 44:115–136] Difficult, highly variable, many hybrids. Not all specimens key easily; sprouts, other extreme forms not included in keys, may require field comparisons. Studies of chromosome numbers, hybridization needed. Inclusion of Salix sessilifolia Nutt. in TJM (1993) based on misidentification of plants belonging to Salix melanopsis. Fruit length as given throughout excludes the stalk (stipe). Hair lengths: minute, < ± 0.5 mm; short, ± 0.5 mm; long, > ± 0.5 mm. Salix commutata Bebb, treated as misapplied to Salix eastwoodiae in TJM (1993), may occur in northern California mountains; Salix bonplandiana expected in s-most California. For alternate treatments, see Dorn (e.g., 2000 Brittonia 52:1–19).
Unabridged references: [Dorn, R. D. 1995. A taxonomic study of Salix section Cordatae subsection Luteae (Salicaceae). Brittonia 47: 160–174. Dorn, R. D. 1998. A taxonomic study of Salix section Longifoliae (Salicaceae) Brittonia 50: 193–210. Dorn, R. D. 2000. A taxonomic study of Salix sections Mexicanae and Viminella subsection Sitchenses (Salicaceae) in North America. Brittonia 52: 1–19.]
Shrub < 4 m. Stem: twigs yellow-green or red-brown, glaucous or not, shaggy-hairy, glabrous in age. Leaf: stipules leaf-like, early-deciduous or not; petiole 1–14 mm; young leaves hairy; mature blade 23–99 mm, narrowly oblong to elliptic, short-acuminate to convex (base rounded to subcordate), entire to finely short-slender-serrate, abaxially generally densely woolly-tomentose or hairs long-soft-shaggy or short-silky, wavy, to ± 0. Inflorescence: blooming with leaves, pistillate 11–51 mm, on leafy shoots 2–12 mm; margin of shoot leaves generally finely short-slender-serrate (or entire or gland-dotted); flower bract brown or black. Staminate flower: stamens 2. Pistillate flower: ovary glabrous or generally silky, stalk 0.2–1.6 mm, style 0.5–1.5 mm.
2n=76. Alpine, subalpine meadows, streams, talus; 1600–3800 m. Klamath Ranges, North Coast Ranges, High Cascade Range, High Sierra Nevada, Modoc Plateau, White and Inyo Mountains; to western Canada, Montana, Wyoming, New Mexico. May–Jul [Online Interchange]
Previous taxon: Salix drummondiana
Next taxon: Salix exigua
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 20 2014
Citation for this treatment: [Author of taxon treatment] 2012. Salix, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=42769, accessed on Apr 20 2014
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