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FAGACEAE OAK FAMILY

John M. Tucker

Shrub, tree, evergreen or not; monoecious. Leaf: simple, alternate, petioled; margin entire to lobed; stipules small, generally deciduous. Staminate inflorescence: catkin or stiff spike, many-flowered. Pistillate inflorescence: 1–few-flowered, generally above staminate inflorescence; involucre bracts many, generally overlapping, flat or cylindric. Staminate flower: calyx generally 4–6-lobed, minute; petals 0; stamens 4–12+. Pistillate flower: calyx generally 6-lobed, minute; petals 0; ovary inferior, style branches generally 3. Fruit: 1 nut subtended, partly enclosed by scaly, cup-like involucre or 1–3 nuts subtended, enclosed by spiny, bur-like involucre; mature years 1–2. Seed: generally 1.
7 genera, ± 900 species: generally northern hemisphere. [Li et al. 2004 Int J Plant Sci 165:311–324] Wood of Quercus critical for pre-20th century ship-building, charcoal for metallurgy; some now supply wood (Fagus, Quercus), cork (Quercus suber), food (Castanea, chestnut). Lithocarpus densiflorus moved to Notholithocarpus. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Li, R.-Q., Chen, Z.-D., Lu, A.-M., Soltis, D. E., Soltis, P. S., & Manos, P. S. 2004. Phylogenetic relationships in Fagales based on DNA sequences from three genomes. Int. J. Plant Sci. 165: 311–324.]

Key to Fagaceae

QUERCUS OAK
Evergreen or not. Leaf: stipules small, generally early-deciduous. Staminate inflorescence: catkins, 1–several, pendent, slender, proximal on twig. Pistillate inflorescence: in upper leaf axils, short-stalked; flower generally 1. Staminate flower: stamens 4–10. Pistillate flower: calyx minute, generally 6-lobed; ovary enclosed by involucre. Fruit: nut 1, partly enclosed by cup-like involucre (cup) with appressed scales (nut and cup = acorn); scales tubercled to not; mature years 1 (on younger stems) or 2 (on older stems).
2n=24.
± 600 species: northern hemisphere, to northern South America, India. (Latin: ancient name for oak) [Manos et al. 1999 Molec Phylogen Evol 12:333–349] Many named hybrids; those (3) treated here form widespread populations; most others occur as single individuals, and some but not all of these are mentioned here, under the first parent treated (alphabetically). Reproduction of many species declining due to habitat degradation or loss as well as disease.
Unabridged references: [Manos, P. S., Doyle, J. J., & Nixon, K. C. 1999. Phylogeny, biogeography, and processes of molecular differentiation of Quercus subgenus Quercus (Fagaceae). Molec Phylogen Evol 12: 333–349.]

Key to Quercus

Q. kelloggii Newb. CALIFORNIA BLACK OAK
NATIVE
Tree < 25 m, deciduous; trunk bark deeply furrowed in age, checkered, dark gray-brown to black. Leaf: (6)9–20 cm; petiole (3)10–40 mm; blade widely elliptic, obovate, or ± round, adaxially glabrous, bright green, abaxially finely tomentose, ± glabrous in age, pale green, tip generally acute, bristled, margin lobes generally 6 per leaf, with 1–4 coarse, generally bristle-tipped teeth. Fruit: cup 16–25 mm wide, 15–25 mm deep, generally cup-shaped, scales thin, not tubercled, glabrous to puberulent; nut 20–35 mm, oblong-ovoid, puberulent, tip generally obtuse, shell woolly inside; mature year 2.
Slopes, valleys, woodland, conifer forest; 30–2660 m. California Floristic Province (except Great Central Valley, South Coast, Channel Islands), Modoc Plateau; Oregon, Baja California. Hybridizes with Quercus agrifolia, Quercus parvula var. parvula, Quercus parvula var. shrevei, Quercus wislizeni (Quercus ×morehus Kellogg). Apr–May [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 24 2014
Citation for this treatment: [Author of taxon treatment] 2013. Quercus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=40652, accessed on Oct 24 2014

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click for enlargement Quercus kelloggii
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1995 Saint Mary's College of California

Geographic subdivisions indicated for the distribution of Quercus kelloggii Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.