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PINACEAE PINE FAMILY

J. Robert Haller, Nancy J. Vivrette, & James R. Griffin, except as noted

Shrub, tree, evergreen; monoecious.
Stem: young crown conic; twig not grooved, resinous, generally persistent.
Leaf: simple, generally alternate, sometimes in bundles or appearing ± 2-ranked, linear or awl-like; base decurrent, woody or not, persistent several yrs.
Pollen cone: generally < 6 cm, not woody, deciduous.
Seed cone: generally woody; bracts, scales generally persistent; scale not peltate, fused to or free from subtending bract.
Seed: 2, on scale base adaxially.
10 genera, 193 species: generally n hemisphere; many of great commercial value, supplying > 1/2 of world's timber. —Scientific Editors: Thomas J. Rosatti, Bruce G. Baldwin.
Unabridged references: [Thieret 1993 FNANM 2:352–398]

Key to Pinaceae

PINUS PINE

J. Robert Haller & Nancy J. Vivrette


Stem: young crown conic, mature often rounded or flat; branches ± whorled in young plants; young bark smooth, mature furrowed; bud ± conic, generally resinous.
Leaf: generally 2.5–35 cm, generally sessile, in bundles of (1)2–5; bundles 1 in axils of alternate, awl-like bracts, base in a sometimes deciduous, scaly sheath of bracts, generally persistent several yrs.
Seed cone: often whorled, generally maturing, opening 2nd yr, persistent on stem or not; stalk 0 or < 16 cm; bract included, fused to scale at least basally, minute; scale tip reflexed, elongated 3–7 cm or often with a rounded or angled, often prickled knob < 3 cm.
Seed: coat hard, woody or not.
2n=24.
94 species: n hemisphere. (Latin: pine) Pinus pinea L., stone pine (leaves 2 per bundle, 10–30 cm; seed cone 8–15 cm, maturing in 3 yrs) cultivated in Eur for over 6000 yrs for edible seeds (pine nuts), reportedly naturalized in SnFrB, n ChI.
Unabridged references: [Millar & Critchfield 1988 Madroño 35:39–53]
Unabridged note: Morphological, genetic study of relationships among Pinus jeffreyi, Pinus ponderosa, and Pinus washoensis indicates that the taxa should be classified as 2 species, Pinus jeffreyi and Pinus ponderosa, and that the latter comprises three vars.: Pinus ponderosa var. pacifica (new taxon), Pinus ponderosa var. ponderosa, and Pinus ponderosa var. washoensis (new combination).

Key to Pinus

P. contorta Loudon
NATIVE

Stem: mature bark scaly, thin; trunk 2–34 m (extremely variable at maturity).
Leaf: 2 per bundle, 2.5–8.6 cm; sheath persistent.
Seed cone: pendent, 2–6 cm, brown; stalk ± 0; scale tip knobs angled, prickle < 6 mm. [Online Interchange]

Previous taxon: Pinus balfouriana subsp. balfouriana
Next taxon: Pinus contorta subsp. bolanderi

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.