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LENTIBULARIACEAE BLADDERWORT FAMILY

Barry A. Rice

Annual, perennial herb, carnivorous, of moist or aquatic habitats.
Stem: caudex or stolon, then often with thread-like branches.
Leaf: simple, in rosette, or simple or dissected, emerging from caudex or stolon, with minute, carnivorous bladders ± throughout.
Inflorescence: raceme or 1-flowered, scapose.
Flower: bisexual; calyx lips 2[4], upper 3-lobed, lower 2-lobed, or lips unlobed; corolla 2-lipped, spurred at base, lower lip flat or arched upward, blocking throat or not; stamens 2, epipetalous; ovary superior, chamber 1, placenta generally free-central; stigma unequally 2-lobed, ± sessile.
Fruit: capsule, round, 2-valved, circumscissile, or irregularly dehiscent.
Seed: generally many, small.
3 genera, 330 species: worldwide, especially tropics. [Rice 2006 Growing Carnivorous Plants. Timber Press] —Scientific Editor: Thomas J. Rosatti.

Key to Lentibulariaceae

PINGUICULA BUTTERWORT
Perennial [( annual)] of moist habitats; carnivorous by sticky or slimy leaves.
Stem: caudex.
Leaf: simple, in rosette, fleshy; margins entire, incurving over trapped prey or not; adaxially with stalked glands that capture small organisms, sessile glands that digest them.
Inflorescence: 1-flowered [rarely 2-flowered]; scape succulent, glandular, bracts 0.
Flower: calyx upper lip 3-lobed, lower lip 2-lobed; corolla lower lip often hairy, generally not blocking throat.
Fruit: 2-valved.
± 92 species: Am, Eur, Medit. (Latin: ± fat, from leaf surface)

P. macroceras Link HORNED BUTTERWORT
NATIVE

Leaf: 2–5 cm, elliptic to ovate, green to dark brown.
Inflorescence: 1–5 per rosette, 1–2 dm.
Flower: corolla (including spur) 13–21 mm, 5-lobed, (pale) blue-violet, center of lower lip white; lobes obovate, throat hairy, spur 6–9 mm.
2n=64. Moist slopes, cliffs, serpentine banks; < 1830 m. n Klamath Ranges (Del Norte, Siskiyou cos.); Oregon to Alaska, Russia; also Japan. [Pinguicula vulgaris subsp. macroceras (Link) Calder & Roy L. Taylor] May not be distinct from Pinguicula vulgaris L. (also 2n=64); plants overwinter as dense bud, proliferate by detachment of basal daughter buds (gemmae). Apr–Jul [Online Interchange] {CNPS list}
Unabridged distribution outside California: sw OR; also n OR to AK, Russia; also Japan.
Unabridged note: May not be distinct from Pinguicula vulgaris L. (also 2n=64); CA, sw OR plants with calyx lobe tips ± blunt, corolla lobes non-overlapping, sometimes treated elsewhere as Pinguicula macroceras subsp. nortensis J. Steiger & J.H. Rondeau (which might not have been effectively published); plants overwinter as dense bud, proliferate by detachment of basal daughter buds (gemmae).

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
View all CCH records

 

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.