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PTERIDACEAE BRAKE FAMILY

Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux

Perennial, in soil or on or among rocks; rhizome creeping to erect, scaly.
Leaf: generally all ± alike (or of 2 kinds, fertile, sterile), generally < 50 cm, often < 25 cm; stipe generally thin, wiry, often dark, ×-section with vascular strands generally 1–3, less often many in circle; blade generally pinnate or ± palmate-pinnate (see Adiantum), often >= 2- compound, abaxially often with glands, ± powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or other, veins generally free.
Sporangia: in sori or not, marginal, submarginal, or along veins, covered by recurved, often modified segment margins (false indusia) or not; true indusia 0; spores spheric, sides flat or not, scar with 3 radiating branches.
± 40 genera, 500 species: worldwide, especially dry areas. [Windham 1993 FNANM 2:122–186] Definition of Cheilanthes, related genera problematic; traditional limits often untenable. —Scientific Editors: Alan R. Smith, Thomas J. Rosatti.

Key to Pteridaceae

NOTHOLAENA
Plant in soil or granite rock crevices; rhizome short-creeping to ± erect, scales lance- linear.
Leaf: stipe generally cylindric, dark brown to black, glabrous to ± scaly; blade 2–4- pinnate, segments generally sessile, ± narrower at base or not.
Sporangia: in ± continuous, marginal bands; segment margin recurved, partly covering sporangia, unmodified; spores finely ridged or granular, often ± black.
34 species: generally Mex, sw US, few in Caribbean, South America. (Greek: false cloak, from leaf blade margin not reflexed as it is in Cheilanthes)
Unabridged references: [Tryon 1956 Contr Gray Herb 179:1–106]

N. californica D.C. Eaton CALIFORNIA CLOAK-FERN
NATIVE
Rhizome scales rigid, with ± black midrib ± to margins, finely ciliate.
Leaf: 3- pinnate, ± 3–13 cm; blade axes brown to black, glabrous or with white to yellow exudate; lowermost pinnae each more developed on basal side; segment abaxially covered with white to yellow exudate, hairs 0, scales 0, adaxially sparsely dotted with white to yellow exudate.
Sporangia: 32-spored.
n=2n=150. Dry rocky slopes, rock crevices, under rock ledges; 200–1300 m. s Channel Islands, San Gabriel Mountains, San Bernardino Mountains, Peninsular Ranges, Desert Mountains, Sonoran Desert; Arizona, nw Mexico. Apogamous. At least 2 entities in CA chemically distinct: 1 with pale to bright yellow exudate on leaf abaxially (Notholaena californica subsp. californica), 1 with white exudate on leaf abaxially (Notholaena californica subsp. leucophylla Windham). Gene flow between them where they overlap geog (s CA) 0 due to apogamy, and pentaploids in AZ suggest further study needed to decide if taxonomic recognition warranted. [Online Interchange]
Unabridged note: There are Consortium records that, if verified, would voucher elevations down to 15 m. The following accessions, if verified, would represent range extensions (as indicated): SBBG19489 (SCo).

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.