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Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux, except as noted

Perennial herb, in soil or on or among rocks; rhizome creeping to erect, scaly. Leaf: generally all ± alike (or of 2 kinds, fertile, sterile), generally < 50 cm, often < 25 cm; stipe generally thin, wiry, often dark, ×-section with vascular strands generally 1–3, less often many in circle; blade generally pinnate or ± palmate-pinnate (see Adiantum), often >= 2-compound, abaxially often with glands, ± powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or other, veins generally free. Sporangia: in sori or not, marginal, submarginal, or along veins, covered by recurved, often modified segment margins (false indusia) or not; true indusia 0; spores spheric, sides flat or not, scar with 3 radiating branches.
± 40 genera, 500 species: worldwide, especially dry areas. [Windham 1993 FNANM 2:122–186] California members of Cheilanthes moved to the distantly related Myriopteris; Pellaea breweri to be moved as well, from a to-be-redefined Pellaea; traditional, often untenable limits of genera outside California also being clarified using molecular phylogenetics. —Scientific Editors: Alan R. Smith, Thomas J. Rosatti.

Key to Pteridaceae

Plant in soil or granite rock crevices; rhizome short-creeping to ± erect, scales lance-linear. Leaf: stipe generally cylindric, dark brown to black, glabrous to ± scaly; blade 2–4-pinnate, segments generally sessile, ± narrower at base or not. Sporangia: in ± continuous, marginal bands; segment margin recurved, partly covering sporangia, unmodified; spores finely ridged or granular, often ± black.
34 species: generally Mexico, southwestern United States, few in Caribbean, South America. (Greek: false cloak, from leaf blade margin not reflexed as it is in Cheilanthes)
Unabridged references: [Tryon 1956 Contr Gray Herb 179:1–106]

N. californica D.C. Eaton CALIFORNIA CLOAK-FERN
Rhizome scales rigid, with ± black midrib ± to margins, finely ciliate. Leaf: 3-pinnate, ± 3–13 cm; blade axes brown to black, glabrous or with white to yellow exudate; lowermost pinnae each more developed on basal side; segment abaxially covered with white to yellow exudate, hairs 0, scales 0, adaxially sparsely dotted with white to yellow exudate. Sporangia: 32-spored.
n=2n=150. Dry rocky slopes, rock crevices, under rock ledges; 200–1300 m. South Coast, s Channel Islands, Transverse Ranges, Peninsular Ranges, Desert Mountains, Sonoran Desert; Arizona, northwestern Mexico. [Notholaena candida (M. Martens & Galeotti) Hook. var. accessita Jeps.] Apogamous. At least 2 entities in California chemically distinct: 1 with pale to bright yellow exudate on leaf abaxially (Notholaena californica subsp. californica), 1 with white exudate on leaf abaxially (Notholaena californica subsp. leucophylla Windham). Gene flow between them where they overlap geographically (s California) 0 due to apogamy, and pentaploids in Arizona suggest further study needed to decide if taxonomic recognition warranted. [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 29 2015
Citation for this treatment: [Author of taxon treatment] 2013. Notholaena, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 29 2015

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Geographic subdivisions indicated for the distribution of Notholaena californica Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.