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HALORAGACEAE WATER-MILFOIL FAMILY

Adolf Ceska & Oldriska Ceska

[Annual, shrub] perennial herb, generally aquatic or semiterrestrial, dioecious or monoecious. Leaf: cauline, opposite, alternate or whorled; submersed blades pinnately divided, segments thread-like; emergent leaves simple, entire to divided. Inflorescence: raceme, spike, or panicle; flowers 1 or clustered, short-pedicelled to ± sessile. Flower: generally unisexual, small; calyx tube short, fused to ovary, lobes 2–4; petals generally 2–4; stamens 4 or 8, filaments generally short; ovary inferior, chambers 1–4, styles 2–4, separate, stigmas generally plumose. Fruit: fleshy or of nut-like mericarps, dehiscent or not. Seed: generally 1 per chamber.
6–8 genera, ± 100 species: especially southern hemisphere, some cultivated. [Aiken & McNeill 1980 J Linn Soc Bot 80:213–222] Haloragis erecta (Murray) Eichler not naturalized in California. —Scientific Editor: Bruce G. Baldwin.

MYRIOPHYLLUM WATER-MILFOIL
Plant from rhizomes, occasionally with overwintering buds (late in growing season); occasionally terrestrial. Stem: simple or branched, generally green. Leaf: submersed leaves generally whorled, 3–6 per node; emergent leaves entire to pinnately divided, occasionally bract-like. Inflorescence: generally emergent, spike-like, simple or branched, terminal, flowers in whorls. Flower: proximal pistillate, middle occasionally bisexual, distal staminate; calyx lobes 4; petals generally 4, ephemeral on staminate flowers, minute or 0 on pistillate flowers; stamens generally 8; ovary 4-chambered. Fruit: mericarps 4, nut-like.
± 60 species: worldwide. (Greek: many leaves, from leaf segments) [Ceska et al. 1986 Brittonia 38:73–81] Myriophyllum specimens best collected in flower or fruit.
Unabridged note: Plants should be washed first and then "floated" on the herbarium mounting paper in a shallow dish with water [see Ceska & Ceska 1987 Ann Missouri Bot Gard 73:825–827].

Key to Myriophyllum

M. verticillatum L. WHORLED WATER-MILFOIL
NATIVE
Monoecious; winter buds club-shaped at ends of non-flowering branches, generally in fall. Stem: > 1 m, dark green to ± black when dry. Leaf: submersed leaves < 5 cm, segments 18–34, < 10 mm, linear, Inflorescence: spike, 5–12 cm, emergent; bracts 3–8(10) mm, pinnately dissected, segments linear.
Lakes, marshes; < 2072 m. North Coast, n&s High Sierra Nevada, Great Basin Floristic Province; to Alaska, Canada, eastern United States, Eurasia. Jul–Sep [Online Interchange]
Unabridged note: Recognizable by stems that are dark green when fresh and ± black when dry, and by inflorescence bracts that resemble miniature leaves. Club-shaped winter buds develop later in season.

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 26 2014
Citation for this treatment: [Author of taxon treatment] 2013. Myriophyllum, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=34279, accessed on Jul 26 2014

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Bioregions in which Myriophyllum verticillatum occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.