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Ronald L. Hartman & Richard K. Rabeler, except as noted

Annual to perennial herb; rarely dioecious (Silene), taprooted or rhizome generally slender. Leaf: simple, generally opposite (subwhorled), entire, pairs at nodes often ± connected at bases; stipules generally 0; petiole generally 0. Inflorescence: generally cyme, generally open; flowers 1–many; involucre generally 0 (present in Dianthus, Petrorhagia). Flower: generally bisexual, radial; hypanthium often present but obscure; sepals (4)5, ± free or fused into a tube, margins generally scarious, more so on inner 2 or not, tube generally not scarious, awns generally 0; petals (4)5 or 0, generally tapered to base (or with claw long, limb expanded), entire to 2–several-lobed, limb generally without scale-like appendages adaxially, generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries 0 or 5; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2–5 with 0 branches or 1 with 2–3 branches. Fruit: capsule or utricle (rarely ± dehiscent), generally sessile. Seed: appendage generally 0 (present in Moehringia).
83 or 89 genera, 3000 species: widespread, especially arctic, alpine, temperate northern hemisphere; some cultivated (Agrostemma, Arenaria, Cerastium, Dianthus, Gypsophila, Lychnis, Sagina, Saponaria, Silene, Vaccaria). [Rabeler & Hartman 2005 FNANM 5:3–215] Apetalous Caryophyllaceae can also be keyed in Rabeler & Hartman 2005 FNANM 5:5–8. —Scientific Editor: Thomas J. Rosatti.

Key to Caryophyllaceae

Perennial herb, ascending to erect, rhizomed. Leaf: not congested at base of flowering stems; petiole short or 0; blade ± lanceolate or elliptic, acute; vein 1. Inflorescence: terminal or axillary; flowers 1–5; pedicels 2–25+ mm. Flower: hypanthium short, obscure; sepals 5, ± free, 2.8–6 mm, ± ovate, minutely ciliate; petals 5, 2–8 mm, entire; stamens on disk; styles 3, 1.5–1.8 mm. Fruit: capsule, widely ovoid; valves 6, ± recurved. Seed: few, red-brown to ± black; appendage ± 0.7 mm, ± elliptic, spongy.
25 species: northern temperate. (P.H.G. Moehring, Danzig naturalist, 1710–1791) [Rabeler & Hartman 2005 FNANM 5:70–72]

M. macrophylla (Hook.) Fenzl LARGE-LEAVED SANDWORT
Stem: simple to branched, 2–18 mm, ± angled or grooved; hairs minute, peg-like; rhizome slender, branched. Leaf: 1.5–5 cm, ± evenly spaced, ± thin; margin smooth to minutely granular, generally ciliate in basal 1/2. Inflorescence: bracts 1–4 mm, margin widely scarious, minutely ciliate; pedicels ascending to erect, ± spreading in fruit or not. Flower: sepals acute to acuminate, margin scarious, midrib ± keeled; petals ± round. Fruit: black. Seed: 1.5–2.2 mm, widely elliptic; tubercles minute, low, rounded.
2n=48. Moist, shaded slopes, rocky ridges, summits, pine, oak forests, serpentine; 300–1800 m. Northwestern California, n&s High Sierra Nevada, San Francisco Bay Area, Outer South Coast Ranges, Peninsular Ranges; to British Columbia, Montana, Colorado, New Mexico, also ± northern Canada, northeastern North America; Asia. [Arenaria macrophylla Hook.] Late spring–summer [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 30 2015
Citation for this treatment: [Author of taxon treatment] 2013. Moehringia, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 30 2015

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click for enlargement Moehringia macrophylla
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1995 Saint Mary's College of California

Geographic subdivisions indicated for the distribution of Moehringia macrophylla Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.