Jepson Herbarium
		University of California, Berkeley

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Annual to shrub [ tree, vine], glabrous to hairy, generally aromatic.
Stem: generally erect, generally 4-angled.
Leaf: generally simple to deeply lobed, generally opposite, generally gland-dotted.
Inflorescence: generally cymes, generally many in dense axillary clusters surrounding stem, generally separated by evident internodes or collectively crowded, spike- or panicle-like, occasionally head-like or raceme, subtended by leaves or bracts; flowers sessile or pedicelled.
Flower: generally bisexual; calyx generally 5-lobed, radial to bilateral; corolla generally bilateral, 1–2-lipped, upper lip entire or 2-lobed, ± flat to hood-like, occasionally 0, lower lip generally 3-lobed; stamens generally 4, epipetalous, generally exserted, paired, pairs generally unequal, occasionally 2, staminodes 2 or 0; ovary superior, generally 4-lobed to base chambers 2, ovules 2 per chamber, style 1, generally arising from center at junction of lobes, stigmas generally 2.
Fruit: generally 4 nutlets, generally ovoid to oblong, smooth.
± 230 genera, 7200 species: worldwide. Many cultivated for herbs, oils (Lavandula, lavender; Mentha, mint; Rosmarinus, rosemary; Thymus, thyme), some cult as ornamental (in CA Cedronella, Leonotis, Monarda, Phlomis). [Harley et al. 2004 Fam Generally Vasc Plant 7:167–275] Moluccella laevis L., shell flower, historical waif in CA. Satureja calamintha (L.) Scheele subsp. ascendens (Jordan) Briq. reported as alien but not naturalized. Salazaria moved to Scutellaria; CA Satureja moved to Clinopodium. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Lamiaceae

Perennial from rhizome, glabrous or hairy.
Stem: erect, branched or not.
Leaf: short-petioled to sessile; blade generally ovate to lanceolate, margin toothed to deeply lobed or cut.
Inflorescence: head-like in leaf axils.
Flower: calyx generally 5-lobed, lobes ± equal, obtuse to short-awned; corolla ± bilateral, not 2-lipped, generally 4-lobed, lobes ± unequal; stamens 2, exserted, staminodes 2, minute, club-shaped; style exserted.
Fruit: nutlets ± compressed, truncate to rounded, edges corky-thickened.
14 species: temperate North America, Eurasia, Australia. (Greek: wolf foot, from French common name) [Moon et al. 2006 J Plant Res 119:633–644]
Unabridged references: [Moon & Hong 2003 Ann Bot Fenn 40:191–198; Henderson 1962 Amer Midl Naturalist 68:95–135]

Rhizomes slender, tips abruptly thicker, tuber-like.
Stem: ascending to erect, 1–5 dm, puberulent to finely strigose.
Leaf: generally short-petioled, 2–6(8) cm; blade elliptic to lanceolate, generally serrate, glabrous to sparsely puberulent.
Flower: calyx lobes ovate, obtuse to acute; corolla 2.5–4 mm, > calyx, white.
Fruit: nutlet 1–2 mm, tip truncate, ± finely toothed.
Moist areas, marshes, near springs; < 2000 m. Northwestern California, Cascade Range, Sierra Nevada, Modoc Plateau;

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.