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SCROPHULARIACEAE FIGWORT FAMILY

Margriet Wetherwax, except as noted

Annual to tree, generally glandular, some ± aquatic. Stem: round to square in ×-section. Leaf: simple, generally alternate, generally ± entire; stipules generally 0 (present in Limosella). Inflorescence: spike to panicle (head-like), generally bracted, or flowers 1–4 in axils. Flower: generally bisexual; calyx lobes 4–5; corolla bilateral to radial, lobes 4–5; stamens epipetalous, 4–5, 5th a staminode in Scrophularia; pistil 1, ovary superior, chambers generally 2, placentas axile, style 1, stigma lobes generally 2. Fruit: capsule, generally ± ovoid, loculicidal or septicidal, or drupe-like.
± 65 genera, 1700 species: ± worldwide; some cultivated as ornamental (Verbascum). [Olmstead et al. 2001 Amer J Bot 88:348–361] Other taxa moved to Plantaginaceae (Antirrhinum, Bacopa, Collinsia, Cymbalaria, Digitalis, Dopatrium, Gambelia, Gratiola, Hebe, Holmgrenanthe, Keckiella, Kickxia, Limnophila, Linaria, Lindernia, Maurandella, Mohavea, Nothochelone, Penstemon, Pseudorontium, Stemodia, Synthyris, Tonella, Veronica), Orobanchaceae (Bellardia, Castilleja, Cordylanthus, Orthocarpus, Parentucellia, Pedicularis, Triphysaria), Phrymaceae (Mimulus). —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Albach et al. 2005 Amer J Bot 92:297–315]

Key to Scrophulariaceae

LIMOSELLA MUDWORT
Annual, stoloned, occasionally partly submersed. Leaf: basal, erect; stipules generally present; petioles present or not. Inflorescence: scapose; flower 1. Flower: calyx bell-shaped, lobes 5, ovate, acute; corolla radial, bell-shaped, lobes 5, adaxially sparsely papillate; stamens 4; style generally ± terminal, stigmas generally fused, head-like. Fruit: elliptic to spheric, chambers 2 proximally, 1 distally. Seed: many, minute.
± 15 species: worldwide. (Latin: mud seat, from habitat)

Key to Limosella

L. australis R. Br. DELTA MUDWORT
NATURALIZED
Tufted. Leaf: 1–3 cm, awl-like, cylindric, obtuse; stipules 0 or tapered; petiole 0. Inflorescence: pedicels < leaves, curved in fruit. Flower: calyx lobes <= tube; corolla ± 3 mm, lobes rounded, white to lavender-blue; stamens attached at 1 level; style > ovary, ± 1 mm, head-like, straight to ± curved, ± terminal. Fruit: spheric.
2n=20. Muddy or sandy intertidal flats, brackish water; < 10 m. Deltaic Great Central Valley (near Antioch), Central Coast (Abbott's Lagoon, Marin Co.); native to eastern coast North America. [Limosella subulata E. Ives] Poorly known in western North America, seeds possibly brought in with ship ballast. Apr [Online Interchange]

Previous taxon: Limosella aquatica
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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 25 2014
Citation for this treatment: [Author of taxon treatment] 2013. Limosella, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=31029, accessed on Oct 25 2014

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click for enlargement Limosella australis
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2008 Louis-M. Landry

Geographic subdivisions indicated for the distribution of Limosella australis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.