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MONTIACEAE MINER'S LETTUCE FAMILY

John M. Miller, except as noted

Annual to perennial herb; generally fleshy. Stem: 1–many, generally glabrous. Leaf: simple, alternate or opposite. Inflorescence: axillary or terminal; cyme, raceme, panicle, umbel, or flower 1. Flower: bisexual, radial; sepals generally 2(9), free; petals (1)2–19, free or ± fused; stamens 1–many, epipetalous or not, anthers pink, rose, or yellow; ovary superior, chamber 1, ovules 1–many, placenta basal or free-central; styles (0)1–8, generally fused at base, branched. Fruit: capsule, circumscissile or 2–3-valved. Seed: 1–many, shiny or ± pebbly or sculptured, black or gray, generally with oil-filled appendage as food for ants.
± 22 genera, ± 230 species: generally temperate America, Asia, Australia, Europe, Kerguelen Is, New Zealand, southern Africa, poorly represented in Europe; some cultivated (Lewisia, Calandrinia). [Ogburn & Edwards 2009 Amer J Bot 96:391–408] Details of flowers, seeds require 20× magnification. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Applequist et al. 2006 Syst Bot 31:310–319; Nyffeler et al. 2008 Haseltonia 14: 26–36]
Unabridged note: May include Hectorellaceae.

Key to Montiaceae

LEWISIA

John M. Miller & Lauramay T. Dempster

Perennial herb generally from short, thick, ± branched taproot; tuberous root generally 0 (or spheric). Stem: prostrate to erect, scape-like or branched. Leaf: generally in basal rosette and cauline, simple, entire or not; base wide; margin generally ± translucent. Inflorescence: ± scapose; cyme, panicle, raceme, or ± umbel; stems 1–many, generally leafless but bracted, disjointing in age or not, 1–many flowered; pedicel 0–30 mm. Flower: sepals 2–8, free, persistent; petals 4–19, white, cream, yellow, orange, pink, rose, purple, overlapped in bud, often with pink or dark purple veins; stamens 1–50; styles 2–8, fused at base, stigmas 2–8, thread-like. Fruit: 6–9 mm, spheric or ovoid, circumscissile near middle or below, translucent. Seed: 1–50, dark, generally shiny, smooth or finely tubercled, 1–4 mm in size.
18 species: western North America, 16 in flora. (Captain Meriwether Lewis, of Lewis & Clark Expedition, 1774–1809) [Wilson et al. 2005 W N Amer Naturalist 65:345–358] Many hybrids, cultivated, including Lewisia ×whiteae Purdy in California; Lewisia columbiana (A. Gray) B.L. Rob. not in California.
Unabridged references: [Colley & Mineo 1985 Pacific Hort 46; Davidson 2000 Lewisias (Portland); Dempster 1996 Madroño 43:415–416; Elliott 1966 Bull Alpine Gard Soc Gr Brit 34:1–76; Foster, Carroll, & Hipkins 1997 Fremontia 25:15–19; Gankin & Hildreth 1968 Four Seasons 2(4):12–14; Heckard & Stebbins 1974 Brittonia 26:305–308; Hershkovitz 1990 Phytologia 68:267–270; Hershkovitz & Hogan 2003 FNANM 4:476–485; Hohn 1975 Ph.D. Dissertation, Univ of Washington; Mathew 1989 Kew Magazine Monogr; Daubenmire 1975 Syesis 8:9–23]
Unabridged note: Many hybrids, cultivars (Mathew 1989), including Lewisia ×whiteae Purdy in California, as well as Lewisia columbiana (Howell ex A. Gray) B.L. Rob. from Douglas Co., Oregon north to British Columbia; Lewisia maguirei A.H. Holmgren endemic to Nevada; Lewisia sacajaweana B.L. Wilson & E. Rey-Vizgirdas restricted to Idaho; Lewisia tweedyi, endemic to northern Cascades of Washington, British Columbia now placed in Cistanthe.

Key to Lewisia

L. oppositifolia (S. Watson) B.L. Rob. OPPOSITE-LEAVED LEWISIA
NATIVE
Leaf: few to many, reduced on lower part of stem or not, 4–11 cm, linear-oblanceolate, entire, tapered to slender base, tip blunt. Inflorescence: stems 1–3, 6–20 cm, each with 2–5-flowered, ± umbel-like cluster; flowers rarely included in leaves; bracts among flowers and below, lanceolate, ± toothed, glands 0; pedicel 20–75 mm. Flower: sepals 2, ± 1/3 × corolla, round or truncate, jagged-dentate, glands 0; petals 8–11, 12–17 mm, ovate, pink fading to white, tips blunt or ± jagged; stamens 8–21; stigmas 3–5. Fruit: 5–6 mm. Seed: 5–15, 1–1.8 mm.
Moist sites, conifer forest; 300–1300 m. Klamath Ranges; southwestern Oregon. Mar–May [Online Interchange] {CNPS list}

Previous taxon: Lewisia nevadensis
Next taxon: Lewisia pygmaea

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 23 2014
Citation for this treatment: [Author of taxon treatment] 2012. Lewisia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=30837, accessed on Apr 23 2014

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click for enlargement Lewisia oppositifolia
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© 2013 Scot Loring

Bioregions in which Lewisia oppositifolia occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.