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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery.
Leaf: generally simple, alternate; generally both basal, cauline; stipules 0.
Inflorescence: generally raceme, generally not bracted.
Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed.
Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not.
Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Medit area, mtns of sw Asia, adjacent c Asia, w North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in c NCoR, Carrichtera annua (L.) DC. in SCo, Iberis sempervirens L., Iberis umbellata L. in PR, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in s NCoRO, CCo. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; CA taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

LEPIDIUM PEPPERGRASS, PEPPERCRESS
Annual to perennial herb ( shrub); hairs 0 or simple.
Leaf: basal rosetted or not, petioled, entire, dentate, to 1–3-pinnately lobed; cauline short-petioled to sessile, base occasionally lobed to clasping.
Inflorescence: elongated or congested.
Flower: sepals erect or spreading, oblong to ovate, base not sac-like; petals linear to obovate, white or yellow (pink or purple), occasionally reduced or 0; stamens 2, 4, or 6.
Fruit: silicle, generally dehiscent, oblong to ovate, obcordate, or round (spectacle-shaped), flat perpendicular to septum (inflated), unsegmented.
Seed: 2(4), gelatinous when wet; wing narrow or 0.
220 species: all continents except Antarctica. (Greek: little scale, from fruit) [Al-Shehbaz et al. 2002 Novon 12:5–11]
Unabridged references: [Hitchcock 1936 Madroño 3:265–300]

Key to Lepidium

L. latipes Hook.
NATIVE
Annual, puberulent or stiff-hairy.
Stem: erect decumbent, 1 or several from base, 0.2–1.5(3.8) dm, simple or branched.
Leaf: basal not rosetted, early- deciduous, 2–10 cm, linear, entire, dentate, or pinnately divided into 2–10 lobe pairs; mid- cauline similar, smaller, entire, base tapered, not lobed.
Inflorescence: compact, cylindric to head-like, elongated or not; rachis puberulent, hairs cylindric.
Flower: sepals 1.1–1.4 mm, ovate; petals 1.9–3 mm, 0.8–1.3 mm wide, obovate- oblong, ± green, hairy abaxially, margin fringed; stamens 4.
Fruit: 5–7 mm, 2.8–4 mm wide, oblong-ovate, tip winged, notch 1.4–2.8 mm; valve walls thick, with a mixture of long, much shorter hairs, strongly net-veined; style 0; pedicel 2.5–5 mm, strongly flattened, appressed, erect to ± ascending, puberulent throughout (only adaxially).
Seed: 2–2.4 mm, oblong.
Alkaline soils, vernal pool margins, salt marsh edges, pastures; < 700 m. North Coast, North Coast Ranges, Great Central Valley, Central Coast, San Francisco Bay Area, Inner South Coast Ranges, South Coast; Baja California. [Lepidium latipes var. heckardii Rollins] Mar–Jun [Online Interchange]
Unabridged note: Lepidium latipes var. heckardii differs from Lepidium latipes var. latipes solely on having stems elongated, simple at base (vs short and branched basally), but these 2 forms grow mixed in most populations. There are Consortium records that, if verified, would voucher elevations up to 914 m. The following (and possibly other) accessions, if verified, would represent range extensions (as indicated): SBBG109095, SBBG109096, SBBG34437, SBBG70830, SBBG73770, SBBG80433, SBBG89981, SBBG90923, SBBG91265, UCR20937, RSA136046, RSA341629, RSA4712, RSA511641, RSA535504, RSA589221, RSA627713, RSA627720, SBBG105811, SBBG70831, SBBG87037, SBBG90930, SBBG97238, UC557765, SBBG112875 (ChI).

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
View all CCH records

 

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.