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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

LEPIDIUM PEPPERGRASS, PEPPERCRESS
Annual to perennial herb (shrub); hairs 0 or simple. Leaf: basal rosetted or not, petioled, entire, dentate, to 1–3-pinnately lobed; cauline short-petioled to sessile, base occasionally lobed to clasping. Inflorescence: elongated or congested. Flower: sepals erect or spreading, oblong to ovate, base not sac-like; petals linear to obovate, white or yellow (pink or purple), occasionally reduced or 0; stamens 2, 4, or 6. Fruit: silicle, generally dehiscent, oblong to ovate, obcordate, or round (spectacle-shaped), flat perpendicular to septum (inflated), unsegmented. Seed: 2(4), gelatinous when wet; wing narrow or 0.
220 species: all continents except Antarctica. (Greek: little scale, from fruit) [Al-Shehbaz et al. 2002 Novon 12:5–11]
Unabridged references: [Hitchcock 1936 Madroño 3:265–300]

Key to Lepidium

L. dictyotum A. Gray
NATIVE
Annual; hairs stiff, spreading. Stem: erect to ascending, 0.2–1.3(2) dm, few to many from base, outer decumbent, simple. Leaf: basal not rosetted, early-deciduous, (1.5)2.2–5.7(7) cm, pinnately lobed or divided, lobes linear to narrowly oblong, entire; mid-cauline sessile, blade 1–5 cm, 0.5–2 mm wide, linear, entire (linear lobed), base tapered, not lobed. Inflorescence: elongate, rachis hairs stiff, cylindric, straight. Flower: sepals 0.7–1.1 mm; petals 0; stamens 4. Fruit: 2.5–3.5 mm, 2–2.8 mm wide, ovate, flat, tip winged, notch 0.5–0.7(0.8) mm, U-shaped; valve walls thin, stiff-hairy (glabrous), strongly net-veined; style 0; pedicel (1.6)1.9–2.5(3) mm, strongly flattened, erect to ± ascending, stiff-hairy throughout (only adaxially). Seed: 1.2–1.8 mm, ovate.
Saline soils, dry stream beds, roadsides, sandy flats, fields, meadows, dried pools; < 1600 m. Inner North Coast Ranges, Sierra Nevada Foothills, Great Central Valley, Inner South Coast Ranges, e South Coast, n Western Transverse Ranges, Great Basin Floristic Province, w Mojave Desert; to Washington, Utah, Baja California. Mar–Jun [Online Interchange]

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Next taxon: Lepidium didymum

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 30 2014
Citation for this treatment: [Author of taxon treatment] 2013. Lepidium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=30532, accessed on Aug 30 2014

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Geographic subdivisions indicated for the distribution of Lepidium dictyotum Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.