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PLANTAGINACEAE PLANTAIN FAMILY

Margriet Wetherwax, except as noted

Annual to shrub, some aquatic. Leaf: basal or cauline, alternate or opposite (whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0. Inflorescence: raceme, spike, or flowers axillary in 1–few-flowered clusters; flowers few to many, each subtended by 1 bract. Flower: unisexual or bisexual, radial or bilateral; sepals 4–5, generally fused at base; corolla 4–5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1–2, anthers opening by 2 slits; ovary superior, [1]2–4-chambered, style 1, stigma lobes 0 or 2. Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
110 genera, ± 2000 species: worldwide, especially temperate. [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2001 Molec Phylogen Evol 16:96–112] Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic California genera of Scrophulariaceae (except Buddleja, Limosella, Mimulus, Myoporum, Scrophularia, Verbascum). California Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.

Key to Plantaginaceae

KECKIELLA BUSH PENSTEMON

Margriet Wetherwax & Noel H. Holmgren

Subshrub or shrub. Stem: wand-like to much-branched. Leaf: drought-deciduous, ± opposite or in 3s, distal occasionally alternate; sessile to short-petioled. Inflorescence: panicle or spike-like; bracts reduced. Flower: calyx lobes 5, ± equal; corolla ± white, ± yellow, or red, short-glandular outside, upper lip ± hooded, 2-lobed, lobes rounded, external in bud, lower lip rounded, lobes often reflexed; stamens 4, filaments attached at 1 level, densely nonglandular-hairy at base, anthers small, glabrous, anther sacs generally spreading flat at dehiscence; staminode well developed, glabrous to densely bearded; nectary a disk; stigma unlobed. Fruit: septicidal and sometimes also loculicidal at tip, ovoid. Seed: many, irregularly angled.
7 species: especially California, western Nevada, western Arizona, Baja California. (David D. Keck, California botanist, 1903–1995; Keckia used earlier, for genus of fossil algae) [Freeman et al. 2003 Syst Bot 28:782–790] Red-flowered species are hummingbird-pollinated.
Unabridged references: [Straw 1966 Brittonia 18:80–95; Wolfe et al. 2002 Syst Bot 27:138–148]

Key to Keckiella

K. lemmonii (A. Gray) Straw
NATIVE
Stem: 5–15 dm; young stems glabrous, glaucous, wand-like. Leaf: opposite; blade 10–65 mm, lanceolate to ovate, base rounded to widely wedge-shaped, margin generally 2–12-toothed. Inflorescence: panicle, glandular-hairy; flowers pedicelled. Flower: calyx 3.5–6.7 mm, lobes ± lanceolate; corolla 11–15 mm, purple-brown, lower lip pale yellow, brown-purple-lined, tube + expanded throat 5.5–9.5 mm, upper lip 2.6–6 mm; anther sacs 0.6–0.9 mm, valves spreading widely; staminode densely yellow-hairy, exserted.
Rocky slopes, conifer and mixed forests, chaparral; < 2100 m. Northwestern California, n Sierra Nevada; Oregon. Jun–Aug [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 25 2014
Citation for this treatment: [Author of taxon treatment] 2013. Keckiella, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=29884, accessed on Nov 25 2014

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click for enlargement Keckiella lemmonii
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2004 Sheli Wingo

Geographic subdivisions indicated for the distribution of Keckiella lemmonii Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.