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Annual to shrub, some aquatic. Leaf: basal or cauline, alternate or opposite (whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0. Inflorescence: raceme, spike, or flowers axillary in 1–few-flowered clusters; flowers few to many, each subtended by 1 bract. Flower: unisexual or bisexual, radial or bilateral; sepals 4–5, generally fused at base; corolla 4–5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1–2, anthers opening by 2 slits; ovary superior, 2–4-chambered, style 1, stigma lobes 0 or 2. Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
110 genera, ± 2000 species: worldwide, especially temperate. [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2001 Molec Phylogen Evol 16:96–112] Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic California genera of Scrophulariaceae (except Buddleja, Limosella, Mimulus, Myoporum, Scrophularia, Verbascum). California Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Key to Plantaginaceae
Subshrub or shrub. Stem: wand-like to much-branched. Leaf: drought-deciduous, ± opposite or in 3s, distal occasionally alternate; sessile to short-petioled. Inflorescence: panicle or spike-like; bracts reduced. Flower: calyx lobes 5, ± equal; corolla ± white, ± yellow, or red, short-glandular outside, upper lip ± hooded, 2-lobed, lobes rounded, external in bud, lower lip rounded, lobes often reflexed; stamens 4, filaments attached at 1 level, densely nonglandular-hairy at base, anthers small, glabrous, anther sacs generally spreading flat at dehiscence; staminode well developed, glabrous to densely bearded; nectary a disk; stigma unlobed. Fruit: septicidal and sometimes also loculicidal at tip, ovoid. Seed: many, irregularly angled.Key to Keckiella
7 species: especially California, western Nevada, western Arizona, Baja California. (David D. Keck, California botanist, 1903–1995; Keckia used earlier, for genus of fossil algae) [Freeman et al. 2003 Syst Bot 28:782–790] Red-flowered species are hummingbird-pollinated.
Unabridged references: [Straw 1966 Brittonia 18:80–95; Wolfe et al. 2002 Syst Bot 27:138–148]
Shrub 3–6 dm, < 1 m wide. Stem: young stems glabrous or hairy. Leaf: opposite; blade 10–35 mm, (ob)lanceolate to narrowly (ob)ovate, base wedge-shaped, margin entire to 3–5-toothed. Inflorescence: pedicels (and calyces) glandular-hairy and densely coarse-hairy. Flower: calyx 6.4–11 mm, lobes ± lanceolate; corolla 22–40 mm, bright pink to red, tube 17–22 mm, throat indistinct, upper lip 9–15 mm; anther sacs 0.9–1 mm; staminode densely yellow-hairy, included.
Rocky slopes in conifer or hardwood forests, (chaparral); 100–2000 m. Northwestern California, Cascade Range Foothills, n Sierra Nevada Foothills, Sacramento Valley (Sutter Buttes), n&c Central Western California. Jun–Oct [Online Interchange]
Unabridged note: Expanded author citation: Keckiella corymbosa (Benth. ex A. DC.) Straw
Previous taxon: Keckiella cordifolia
Next taxon: Keckiella lemmonii
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Mar 7 2014
Citation for this treatment: [Author of taxon treatment] 2012. Keckiella, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=29883, accessed on Mar 7 2014
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© 2012 Aaron Arthur
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