Jepson Herbarium
		University of California, Berkeley

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Perennial from caudex or rhizome, generally ± hairy.
Stem: often ± leafy on proximal 1/2, rarely trailing and leafy throughout.
Leaf: generally simple, basal and/or cauline, generally alternate, generally petioled; veins ± palmate.
Inflorescence: raceme or panicle, generally ± scapose.
Flower: generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens 3, [4], 5, 8, or 10; pistils 1 ( carpels ± fused, ovary lobed, chambers 1 or 2, placentas generally 2(3), axile or parietal or occasionally proximally axile and distally marginal in ovary lobes) or 2 ( carpels free, placentas marginal), ovary nearly superior to inferior, occasionally more superior in fruit, styles generally 2(3).
Fruit: capsule (generally 2(3)-beaked, valves generally 2(3), generally equal) or 2 follicles.
Seed: generally many, small.
± 30 genera, 600 species: especially n temperate, arctic, alpine; some cultivated (Bergenia, Darmera, Heuchera, Saxifraga, Tellima, Tolmiea). [Soltis et al. 2001 Ann Missouri Bot Gard 88:669–693; Okuyama et al. 2008 Molec Phylogen Evol 46:560–575] CA Mitella moved to Mitellastra, Ozomelis, Pectiantia; Suksdorfia ranunculifolia to Hemieva. Parnassia moved to Parnassiaceae. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Soltis 1988 Syst Bot 13:64–72]

Key to Saxifragaceae

Caudex corm-like, flattened or ovoid to spheric, branched or not, without scales or bulblets.
Leaf: generally 1–3, basal; blade ± round, base cordate to reniform, lobes and teeth shallow.
Inflorescence: cyme, appearing before leaves, glandular; bracts scale-like.
Flower: hypanthium free of ovary, generally truncate at base; petals 5, short-clawed; stamens 10; pistil 1, ovary superior, chambers 2, placentas 2, axile.
Fruit: capsule.
3 species: CA. (W.L. Jepson, CA botanist, 1867–1946) [Ornduff 1969 Brittonia 21:286–298] Heterostylous.

Caudex generally flattened, branched.
Leaf: generally 2–3.
Inflorescence: flowers generally 4–17; peduncle 5–23 cm, red-pink, often drying tan.
Flower: hypanthium 1.5–3 mm, generally > calyx lobes, truncate to tapered at base; calyx lobes 1.3–2 mm, pink; petals generally 3.5–6 mm, persistent, withering, veins deep pink.
Fruit: generally ± green or rose, ± red-striped.
Seed: pale brown.
2n=14. Crevices, especially in slate-like rock; dry, rocky slopes; < 700 m. n&c Sierra Nevada Foothills (El Dorado to Mariposa cos.), n San Joaquin Valley (ne Stanislaus Co.).

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.