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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

ISATIS
[Annual] biennial [perennial herb]; hairs simple or 0. Leaf: rosetted or not, petioled, entire or lobed; cauline sessile, base lobed to clasping. Inflorescence: elongated. Flower: sepals erect to ascending, base not sac-like; petals yellow or cream, not clawed. Fruit: indehiscent, pendent, oblong to oblanceolate or round to ovate, flat perpendicular to septum, unsegmented, winged; style 0, stigma entire. Seed: 1(2), oblong, plump.
± 60 species: Eurasia. (Greek: plant with dark dye)

I. tinctoria L. WOAD
NATURALIZED
Biennial, glaucous, glabrous or hairy proximally. Stem: (3)4–10(15) dm, branched distally. Leaf: basal (3)6–20(25) cm, oblong to oblanceolate, entire to wavy or dentate; cauline entire. Inflorescence: panicle of racemes. Flower: sepals 1.5–2.8 mm; petals 2.5–4 mm, 0.9–1.5 mm wide, oblanceolate, yellow. Fruit: (0.9)1.1–2(2.7) cm, 3–6(10) mm wide, oblong-oblanceolate to elliptic-obovate; pedicel reflexed, 5–10 mm, club-shaped near tip. Seed: 2.3–3.5(4.5) mm, oblong.
2n=14,28. Disturbed areas, pastures; 100–2200 m. Klamath Ranges, Cascade Range, n High Sierra Nevada, Modoc Plateau; western and eastern North America, native to Eurasia. Once cultivated as source of the blue dye woad. Apr–Jul {Noxious weed} [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 23 2014
Citation for this treatment: [Author of taxon treatment] 2013. Isatis, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=29342, accessed on Sep 23 2014

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click for enlargement Isatis tinctoria
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© 2001 CDFA

Geographic subdivisions indicated for the distribution of Isatis tinctoria Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.